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Borophagina

Mammalia - Carnivora - Canidae

Synonymy list
YearName and author
1999Borophagina Wang et al. p. 339
2004Borophagina Wang et al. p. 451
2008Borophagina Wang et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
—Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
—Tetrapoda
—Reptiliomorpha
—Anthracosauria
subclassAmphibiosauriaKuhn 1967
—Cotylosauria()
—Amniota
subclassSynapsida
—Therapsida()
infraorderCynodontia()
—Mammaliamorpha
RankNameAuthor
—Mammaliaformes
classMammalia
—Cladotheria
—Zatheria
subclassTribosphenida()
subclassTheria
—Eutheria()
—Placentalia
—Boreoeutheria
—Laurasiatheria
—Scrotifera
—Ferae()
—CarnivoramorphaWyss and Flynn 1993
—CarnivoraformesFlynn et al.
orderCarnivora
familyCanidae
subfamilyBorophaginae
tribeBorophaginiSimpson 1945
subtribeBorophagina
subtribeBorophagina

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subtr. †Borophagina Wang et al. 1999
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G. †Carpocyon Webb 1969
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†Carpocyon compressus Cope 1890
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Invalid names: Cynodesmus cuspidatus Thorpe 1922 [synonym]
†Carpocyon limosus Webb 1969
†Carpocyon robustus Green 1948
†Carpocyon webbi Wang et al. 1999
G. †Epicyon Leidy 1858
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†Aelurodon haydeni validus Matthew and Cook 1909
†Aelurodon saevus secundus Matthew and Cook 1909
†Epicyon aelurodontoides Wang et al. 1999
†Epicyon haydeni Leidy 1858
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Invalid names: Aelurodon aphobus Merriam 1919 [synonym], Osteoborus ricardoensis Stirton and VanderHoof 1933 [synonym], Osteoborus validus Stirton and Vanderhoof 1933 [synonym], Tephrocyon mortifer Cook 1914 [synonym]
†Epicyon saevus Leidy 1858
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Invalid names: Aelurodon inflatus VanderHoof and Gregory 1940 [synonym]
G. †Paratomarctus Wang et al. 1999
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†Paratomarctus euthos McGrew 1935
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Invalid names: Canis anceps Scott 1893 [synonym]
G. †Protepicyon Wang et al. 1999
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†Protepicyon raki Wang et al. 1999
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: phosphaticsubp
Environment: terrestrialsubc
Locomotion: actively mobilec
Life habit: ground dwellingf
Diet: carnivoref
Diet 2: omnivoref
Reproduction: viviparoussubc
Created: 2005-06-08 10:11:09
Modified: 2005-08-22 19:28:06
Source: f = family, subc = subclass, c = class, subp = subphylum
References: Carroll 1988, Hendy et al. 2009, Nowak 1991, Ji et al. 2002, Lillegraven 1979

Age range: base of the Barstovian to the top of the Clarendonian or 16.30000 to 9.40000 Ma

Collections (215 total)


Time interval Ma Country or state Original ID and collection number
Miocene23.03 - 5.333USA (South Dakota) Tomarctus euthos (18113)
Miocene23.03 - 5.333USA (Nevada) Epicyon saevus (18861)
Early/Lower Hemingfordian20.43 - 15.97USA (Texas) Paratomarctus temerarius (18642)
Hemingfordian20.43 - 15.97USA (Oregon) Paratomarctus temerarius (212623)
Barstovian16.3 - 12.5USA (Montana) Canis anceps (18894)
Barstovian16.3 - 12.5USA (California) Epicyon sp. (19458) Paratomarctus temerarius (19362 19364 19366 19422 19522 19542 19556 19568 214888) Paratomarctus temerarius, Epicyon sp. (19432 19457) Paratomarctus temerarius, Protepicyon raki (19463 19548) Tephrocyon temerarius (19157)
Barstovian16.3 - 12.5USA (Nevada) Carpocyon compressus, Cynodesmus kelloggi (19088)
Barstovian16.3 - 12.5USA (Kansas) Carpocyon compressus (18373)
Barstovian16.3 - 12.5USA (Oregon) Tomarctus brevirostris (19056)
Barstovian16.3 - 12.5Canada (Saskatchewan) Aelurodon saevus (18758 18761 18769 18773 18785 18797)
Barstovian16.3 - 12.5USA (Texas) Carnivora indet. (18627)
Barstovian16.3 - 12.5USA (Colorado) Carpocyon compressus (18203)
Barstovian16.3 - 12.5USA (New Mexico) Epicyon haydeni, Epicyon saevus, Paratomarctus temerarius, Carpocyon sp. (19535) Paratomarctus temerarius (19274 19435 19520) Paratomarctus temerarius, Carpocyon sp. (19287) Tomarctus temerarius (19519)
Barstovian16.3 - 12.5USA (Nebraska) Carpocyon compressus (18132 18173 18205 18285 18286) Carpocyon compressus, Paratomarctus temerarius (18123 18124) Carpocyon cuspidatus (18209) Carpocyon cuspidatus, Tomarctus temerarius (18192 18284 18442) Carpocyon sp. (18206 27481) Paratomarctus temerarius (18109 18131 18144 18155 18172 18265 18272 18281) Tomarctus temerarius (18164 18261 18275)
Early/Lower Barstovian16.3 - 12.5USA (Montana) Tomarctus temerarius (18803)
Early/Lower Barstovian16.3 - 12.5USA (Texas) Paratomarctus temerarius (18665)
Barstovian - Clarendonian16.3 - 9.4USA (Nebraska) Canis haydeni, Canis saevus, Canis temerarius (190543) Carpocyon compressus (18070 18112 18270) Carpocyon webbi, Tomarctus euthos (18077) Paratomarctus euthos (18150 18306)
Barstovian - Hemphillian16.3 - 4.9Mexico (Baja California Sur) Aelurodon inflatus (18731)
Langhian15.97 - 13.82USA (California) Paratomarctus temerarius (59059)
Clarendonian12.5 - 9.4USA (Florida) Epicyon haydeni (18588)
Clarendonian12.5 - 9.4USA (Oklahoma) Epicyon sp. (18130)
Clarendonian12.5 - 9.4USA (California) Aelurodon haydeni, Osteoborus diabloensis (19391) Epicyon aphobus (19322) Epicyon aphobus, Tomarctus robustus (19517) Epicyon saevus (19383 19384 189022)
Clarendonian12.5 - 9.4USA (South Dakota) Aelurodon inflatus (18063) Carpocyon robustus, Epicyon saevus, Paratomarctus euthos (18200) Tomarctus euthos, Aelurodon saevus (18268)
Clarendonian12.5 - 9.4USA (Texas) Carpocyon robustus, Epicyon haydeni, Epicyon saevus, Tomarctus euthos (18355) Epicyon saevus, Paratomarctus euthos (18244) Tomarctus euthos (18283)
Clarendonian12.5 - 9.4USA (Nevada) Aelurodon haydeni (18888) Epicyon saevus (18987 19035) Osteoborus diabloensis (18835)
Clarendonian12.5 - 9.4USA (Colorado) Carpocyon robustus (18986)
Clarendonian12.5 - 9.4USA (Nebraska) Aelurodon mortifer (18017) Aelurodon sp. (18197) Carpocyon robustus (18048 18053 18321) Carpocyon robustus, Epicyon haydeni, Epicyon saevus (18447) Carpocyon robustus, Epicyon haydeni, Epicyon saevus, Paratomarctus euthos (18227) Carpocyon robustus, Epicyon saevus, Paratomarctus euthos (18458) Carpocyon webbi (18079 18148 18260 18419 18466) Carpocyon webbi, Epicyon haydeni, Epicyon saevus (188779) Carpocyon webbi, Paratomarctus euthos (18217 18242) Epicyon haydeni (18142 18218 18233 18277 18430) Epicyon haydeni, Aelurodon sp. (18222) Epicyon haydeni, Epicyon saevus (18214 18468) Epicyon haydeni, Epicyon saevus, Paratomarctus euthos (18135 18496) Epicyon saevus (18024 18059 18118 18171 18196 18202 18221 18246 18251 18301 18309 18335 18336 18374 18378 18398 18446) Epicyon saevus, Aelurodon mortifer (18299) Epicyon saevus, Paratomarctus euthos (18042 18095 18245) Paratomarctus euthos (18056 18291 18302 18333 18425) Tomarctus euthos (18176)
Clarendonian12.5 - 9.4USA (Oregon) Aelurodon sp. (18816 18826) Epicyon saevus (19093) Osteoborus sp. (18819)
Clarendonian12.5 - 9.4USA (Arizona) Carpocyon robustus, Epicyon haydeni (19436) Epicyon haydeni (19315 19429 19483 19543)
Late/Upper Clarendonian12.5 - 9.4USA (Florida) Aelurodon haydeni, Aelurodon saevus (18554)
Late/Upper Clarendonian12.5 - 9.4USA (Nebraska) Epicyon haydeni, Epicyon saevus (18327) Epicyon saevus (18066)
Late/Upper Clarendonian12.5 - 9.4USA (New Mexico) Epicyon sp. (19528)
Late/Upper Clarendonian12.5 - 9.4USA (California) Osteoborus ricardoensis (188947)
Late/Upper Clarendonian12.5 - 9.4USA (Oregon) Aelurodon sp. (19036)
Clarendonian - Hemphillian12.5 - 4.9USA (Wyoming) Tomarctus compressus (18424)
Clarendonian - Hemphillian12.5 - 4.9USA (South Dakota) Epicyon saevus (18352) Epicyon saevus, Paratomarctus euthos (18087)
Clarendonian - Hemphillian12.5 - 4.9USA (Nebraska) Epicyon haydeni (18236) Epicyon haydeni, Epicyon saevus (18295) Epicyon saevus (18069 18256) Epicyon saevus, Paratomarctus euthos (18134) Epicyon validus (18326)
Clarendonian - Hemphillian12.5 - 4.9USA (California) Epicyon saevus (19552)
Clarendonian - Hemphillian12.5 - 4.9USA (Texas) Epicyon saevus (18358)
Tortonian11.62 - 7.246USA (California) Epicyon haydeni (19466)
Late/Upper Miocene11.608 - 5.333USA (Montana) Epicyon haydeni (19009)
Late/Upper Miocene11.608 - 5.333USA (New Mexico) Aelurodon haydeni (19424)
Hemphillian10.3 - 4.9USA (Nebraska) Aelurodon validus, Epicyon saevus (18033) Epicyon haydeni, Epicyon saevus (18323) Epicyon saevus (18258 18431) Epicyon sp. (18158) Epicyon validus (18086 18225)
Hemphillian10.3 - 4.9USA (Kansas) Epicyon aelurodontoides (18403) Epicyon haydeni (18111 18365 18444 18497) Epicyon saevus, Epicyon validus (18415) Epicyon sp. (18046 18263) Epicyon validus (18055)
Hemphillian10.3 - 4.9USA (Alabama) Borophaginae indet. (18559)
Hemphillian10.3 - 4.9USA (Nevada) Epicyon saevus (19095)
Hemphillian10.3 - 4.9USA (Florida) Epicyon haydeni (18514 18527) Epicyon saevus, Osteoborus validus (18560) Osteoborus validus (18563)
Hemphillian10.3 - 4.9USA (California) Osteoborus diabloensis (19551)
Hemphillian10.3 - 4.9USA (Arizona) Aelurodon validus (18933)
Hemphillian10.3 - 4.9USA (New Mexico) Aelurodon haydeni (19288) Epicyon haydeni (19348) Epicyon sp., Carpocyon webbi (19186)
Hemphillian10.3 - 4.9USA (Oklahoma) Carpocyon limosus, Epicyon validus, Epicyon mortifer (18034) Epicyon haydeni (18169 18364) Epicyon saevus (18293)
Hemphillian10.3 - 4.9USA (Texas) Epicyon haydeni, Epicyon saevus (18308) Epicyon saevus, Epicyon validus (18072) Epicyon validus (18198 18211)
Hemphillian10.3 - 4.9USA (Colorado) Epicyon validus (18495)
Hemphillian10.3 - 4.9USA (Idaho) Epicyon haydeni (19085)
Late/Upper Hemphillian10.3 - 4.9USA (Nebraska) Carpocyon limosus (35430)
Late/Upper Hemphillian10.3 - 4.9USA (Florida) Carpocyon limosus (18573) Epicyon haydeni (18532)
early Early Hemphillian10.3 - 4.9USA (Texas) Borophaginae indet. (19538)
early Early Hemphillian - late Early Hemphillian10.3 - 4.9USA (Florida) Epicyon haydeni (18539)