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Itemirus medullaris
Taxonomy
Itemirus medullaris was named by Kurzanov (1976). Its type specimen is PIN 327/699. Its type locality is Dzharakuduk II (general), which is in a Turonian/Turonian delta plain sandstone in the Bissekty Formation of Uzbekistan. It was considered monophyletic by Currie (2000).
Synonymy list
| Year | Name and author |
|---|---|
| 1976 | Itemirus medullaris Kurzanov p. 363 figs. 1-3 |
| 1981 | Itemirus medullaris Nessov p. 91 |
| 1991 | Itemirus medullaris Molnar p. 167 |
| 1997 | Itemirus medullaris Ryan p. 443 |
| 1998 | Itemirus medullaris Makovicky and Norell p. 13 |
| 2000 | Itemirus medullaris Currie p. 450 |
| 2004 | Itemirus medullaris Holtz, Jr. p. 114 |
| 2009 | Itemirus medularis Bever and Norell p. 22 |
| 2009 | Itemirus medullaris Longrich and Currie p. supplementary |
| 2012 | Itemirus medullaris Averianov and Sues p. 284 |
| 2014 | Itemirus medullaris Sues and Averianov p. 227 |
| 2021 | Itemirus medullaris Averianov and Lopatin p. 573 |
| 2021 | Itemirus medullaris Tanaka et al. p. 7 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
†Itemirus medullaris Kurzanov 1976
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Diagnosis
| Reference | Diagnosis | |
|---|---|---|
| H.-D. Sues and A. Averianov 2014 | None of the features in the original diagnosis offered by Kurzanov (1976) has diagnostic value, and two characters cited by this author (position of foramen for endolymphatic duct and presence of “recessus interacusticus”) are based on incorrect anatomical interpretations. Itemirus is referable to Dromaeosauridae based on the following synapomorphies for this clade hypothesized by Turner et al. (2012): 17(0), accessory tympanic recess dorsal to crista interfenestralis absent; 96(0), anterior cervical centra level with or shorter than posterior extent of neural arch; 103(1), parapophyses of posterior trunk vertebrae distinctly projected on pedicels. Within Dromaeosauridae Itemirus medullaris is referable to Dromaeosaurinae on the basis of the following synapomorphies: 91(1), premaxillary tooth crowns asymmetrical (D-shaped) in cross-section, with flat lingual surface; 238(1), jugal process of maxilla ventral to external antorbital fenestra dorsoventrally wide. Among Dromaeosaurinae, Itemirus medullaris differs from Dromaeosaurus albertensis in characters 15(1), basipterygoid recesses on dorsolateral surfaces of basipterygoid processes present; 247(1), anterior denticles of teeth significantly smaller than posterior denticles. Itemirus differs from Utahraptor ostrommaysorum in character 265(1), pleurocoels present in anterior dorsals. Itemirus medullaris differs from both Utahraptor ostrommaysorum and Achillobator giganticus in character 318(1), lateral surfaces of centra of thoracic vertebrae have deep, emarginated fossae. |
Measurements
No measurements are available
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| Source: f = family, o = order | |||||
| References: Marsh 1875, Holtz et al. 2000 | |||||
Age range: base of the Middle Turonian to the top of the Late/Upper Turonian or 93.90000 to 89.80000 Ma
Collections (5 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Middle Turonian - Late/Upper Turonian | Uzbekistan (Navoi) | Itemirus medullaris (46959 58159 58166 58168 type locality: 75943) |