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Pakicetus
Taxonomy
Pakicetus was named by Gingerich and Russell (1981). It is the type genus of Pakicetidae. It was considered monophyletic by Cooper et al. (2009).
It was assigned to Protocetidae by Gingerich and Russell (1981), Carroll (1988), Benton (1993); to Pakicetinae by Gingerich and Russell (1990), McKenna and Bell (1997); to Pakicetidae by Thewissen et al. (1996), Thewissen and Hussain (1998), Thewissen et al. (2001), Thewissen et al. (2001), Geisler and Sanders (2003), McLeod and Barnes (2008), Cooper et al. (2009), Uhen (2010), Gingerich (2012); and to Pakicetidae by van Vliet (2004), Clementz et al. (2006), Gingerich and Cappetta (2014), Gao and Ni (2015), Marx et al. (2016), Berta (2017).
It was assigned to Protocetidae by Gingerich and Russell (1981), Carroll (1988), Benton (1993); to Pakicetinae by Gingerich and Russell (1990), McKenna and Bell (1997); to Pakicetidae by Thewissen et al. (1996), Thewissen and Hussain (1998), Thewissen et al. (2001), Thewissen et al. (2001), Geisler and Sanders (2003), McLeod and Barnes (2008), Cooper et al. (2009), Uhen (2010), Gingerich (2012); and to Pakicetidae by van Vliet (2004), Clementz et al. (2006), Gingerich and Cappetta (2014), Gao and Ni (2015), Marx et al. (2016), Berta (2017).
Species
Synonymy list
Year | Name and author |
---|---|
1981 | Pakicetus Gingerich and Russell |
1988 | Pakicetus Carroll |
1990 | Pakicetus Gingerich and Russell p. 17 |
1993 | Pakicetus Benton p. 761 |
1996 | Pakicetus Thewissen et al. p. 71 figs. Figure 31 |
1997 | Pakicetus McKenna and Bell p. 370 |
1998 | Pakicetus Thewissen and Hussain p. 231 |
2001 | Pakicetus Thewissen et al. p. 277 |
2003 | Pakicetus Geisler and Sanders p. 27 |
2004 | Pakicetus van Vliet p. 143 |
2006 | Pakicetus Clementz et al. p. 359 |
2008 | Pakicetus McLeod and Barnes p. 92 |
2009 | Pakicetus Cooper et al. p. 1291 |
2010 | Pakicetus Uhen p. 199 |
2012 | Pakicetus Gingerich p. 313 figs. Figure 4 |
2014 | Pakicetus Gingerich and Cappetta p. 128 figs. Figure 16 |
2015 | Pakicetus Gao and Ni p. 156 figs. Table 1 |
2016 | Pakicetus Marx et al. p. 98 |
2017 | Pakicetus Berta p. 159 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
G. †Pakicetus Gingerich and Russell 1981
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†Pakicetus attocki West 1980
†Pakicetus calcis Cooper et al. 2009
†Pakicetus chittas Cooper et al. 2009
†Pakicetus inachus Gingerich and Russell 1981
Diagnosis
Reference | Diagnosis | |
---|---|---|
L. N. Cooper et al. 2009 | P4 taller than M1; protocone lobe of P4 prominent; M1 labiolingually narrower than P4; small angle be- tween M2 protocone and labial base of trigon; protocone round- ed, slightly bulbous; M3 trigon base mediolaterally wide, steep anterior and posterior borders; p4 taller than m1-m3, anteropos- teriorly elongated at base. Differs from Nalacetus (H-GSP 18521, 96055) and Ichthyolestes (IPHG 1956 II 7) in: protocone rounded, bulbous. Differs from Nalacetus in: M3 trigon labiolin- gually wider base, more lingually oriented paracone tip. |
Measurements
part | N | mean | minimum | maximum | median | s.d. | |
---|---|---|---|---|---|---|---|
P1 height | 1 | 5.40 | - | - | - | - | |
P2 height | 1 | 16.5 | - | - | - | - | |
P3 height | 1 | 18.0 | - | - | - | - | |
M1 height | 1 | 14.8 | - | - | - | - | |
M2 height | 2 | 12.8 | 12.5 | 13.1 | 12.8 | 0.424 | |
M3 height | 1 | 12.5 | - | - | - | - | |
p2 height | 1 | 20.6 | - | - | - | - | |
p3 height | 1 | 20.6 | - | - | - | - | |
p4 height | 1 | 23.0 | - | - | - | - | |
m1 height | 1 | 14.2 | - | - | - | - | |
m3 height | 1 | 12.0 | - | - | - | - | |
C1 height | 1 | 23.4 | - | - | - | - | |
I1 height | 2 | 17.8 | 17.2 | 18.5 | 17.9 | 0.919 | |
c1 height | 2 | 26.6 | 24.4 | 29.0 | 26.7 | 3.25 |
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Source: subf = subfamily, o = order | |||||
References: Uhen 2004, Thewissen et al. 2001 |
Collections (6 total)
Time interval | Ma | Country or state | Original ID and collection number |
---|---|---|---|
Ypresian | India (Jammu and Kashmir) | Ichthyolestes pinfoldi (45588) | |
Lutetian | Pakistan (North-West Frontier) | P. inachus (28978) | |
Lutetian | Pakistan | P. calcis, P. chittas (93595) | |
Lutetian | Pakistan (Punjab) | Basilosauridae indet. (49002) P. attocki (45529) P. inachus, Protocetus attocki (45586) |