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Zuul crurivastator

Reptilia - Ankylosauridae

Taxonomy
Zuul crurivastator was named by Arbour and Evans (2017) [Differs from all ankylosaurids in the possession of the following autapomorphies: imbricated, peaked frontonasal and frontoparietal caputegulae; prominent longitudinal furrows on the lateral surface of the squamosal horn; lateral caudal osteoderms along the tail club (excluding the knob osteoderms) are strongly concave on the leading edge with posteriorly offset apices; tail club knob dorsoventrally flat, with height to length ratio less than 0.20. Zuul can be further differentiated from other ankylosaurins in the following traits: possesses pyramidal prefrontal, frontoparietal and middle supraorbital caputegulae (unlike the conical caputegulae in Nodocephalosaurus and Talarurus); squamosal horns extend posteriorly well past the nuchal shelf, similar to Scolosaurus but unlike Anodontosaurus, Euoplocephalus, or Ziapelta; postocular caputegulae small and sparsely distributed, similar to Scolosaurus but unlike Anodontosaurus, Euoplocephalus or Ziapelta. Caudal osteoderms lateral to tail club handle are proportionately larger and more sharply pointed than in ankylosaurines from the Nemegt Formation of Mongolia.]. Its type specimen is ROM 75860, a partial skeleton, and it is a 3D body fossil. Its type locality is Theropoda Exp LLC Locality LC02(Q), which is in a Campanian terrestrial horizon in the Judith River Formation of Montana.

Synonymy list
YearName and author
2017Zuul crurivastator Arbour and Evans p. 7 figs. 1-7
2017Zuul crurivastator Arbour and Mallon p. 771
2018Zuul crurivastator Penkalski p. 267

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
Sauropsida
classReptilia
subclassEureptilia()
RankNameAuthor
Romeriida
Diapsida()
Archosauromorpha(Huene 1946)
Crocopoda
ArchosauriformesGauthier 1986
Eucrocopoda
Archosauria()
informalAvemetatarsalia
Ornithodira
Dinosauromorpha
Dinosauriformes
Dinosauria()
Ornithischia()
Genasauria
Thyreophora()
Ankylosauria()
familyAnkylosauridae
subfamilyAnkylosaurinaeNopcsa 1918
genusZuul
speciescrurivastator

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Zuul crurivastator Arbour and Evans 2017
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Diagnosis
ReferenceDiagnosis
P. Penkalski 2018Differs from Euoplocephalus, Dyoplosaurus, Scolosaurus, Anodontosaurus, Oohkotokia, and Platypelta in having many cranial caputegulae with prominent peaks or node-like apices and thickened, raised caudal borders and in having a hemi-concentric, ‘shield’-like arrangement to the frontal caputegulae.
Measurements
No measurements are available
Composition: hydroxyapatiteo
Entire body: yeso
Adult length: 10 to < 100o
Adult width: 1.0 to < 10o
Adult height: 1.0 to < 10o
Architecture: compact or denseo
Ontogeny: accretion, modification of partso
Grouping: gregariouso
Environment: terrestrialo
Locomotion: actively mobileo
Life habit: ground dwellingo
Diet: herbivoreo
Reproduction: oviparouso
Dispersal: direct/internalo
Dispersal 2: mobileo
Created: 2004-12-14 12:50:52
Modified: 2004-12-14 14:50:52
Source: o = order
Reference: Marsh 1875

Age range: Late/Upper Campanian or 83.60000 to 72.20000 Ma

Collections: one only


Time interval Ma Country or state Original ID and collection number
Late/Upper Campanian83.6 - 72.2USA (Montana) Zuul crurivastator (type locality: 190394)