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Ommatinae
Taxonomy
Ommatinae was named by Sharp and Muir (1912). It is extant. Its type is Omma.
It was considered an invalid subgroup of Ommatidae by Tan et al. (2012).
It was assigned to Cupedidae by Ponomarenko (1966), Hong and Wang (1990), Soriano and Delclos (2006), Jarzembowski et al. (2013), Ponomarenko (2014), Jarzembowski et al. (2016), Jarzembowski et al. (2016), Jarzembowski and Wang (2016), Jarzembowski et al. (2017), Jarzembowski et al. (2019), Kirejtshuk (2020); and to Ommatidae by Lawrence and Newton (1995), Tan and Ren (2009), Bouchard et al. (2011), Yamamoto (2017), Cai and Huang (2017), Tihelka et al. (2019), Cai et al. (2022).
It was considered an invalid subgroup of Ommatidae by Tan et al. (2012).
It was assigned to Cupedidae by Ponomarenko (1966), Hong and Wang (1990), Soriano and Delclos (2006), Jarzembowski et al. (2013), Ponomarenko (2014), Jarzembowski et al. (2016), Jarzembowski et al. (2016), Jarzembowski and Wang (2016), Jarzembowski et al. (2017), Jarzembowski et al. (2019), Kirejtshuk (2020); and to Ommatidae by Lawrence and Newton (1995), Tan and Ren (2009), Bouchard et al. (2011), Yamamoto (2017), Cai and Huang (2017), Tihelka et al. (2019), Cai et al. (2022).
Subtaxa
Allophalerus, Blapsium (syn. Blapsidium), Brochocoleus, Burmocoleus, Cionocoleus, Clessidrommatini, Diluticupes, Echinocups, Eurydictyon, Jarzembowskiops, Liassocupes, Lithocupedini, Lithocupes, Monticupes, Notocupoides, Odontomma, Ommatini, Paraodontomma, Pareuryomma (syn. Euryomma), Polyakius, Pronotocupedini, Rhopalomma, Stegocoleus, Tetraphalerini, Tetraphalerites, Tetraphalerus, Tetraphalerus brevicapitis
Synonyms
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Synonymy list
| Year | Name and author |
|---|---|
| 1912 | Ommatinae Sharp and Muir |
| 1962 | Tetraphalerini Crowson |
| 1966 | Notocupedinae Ponomarenko |
| 1966 | Ommatinae Ponomarenko |
| 1969 | Lithocupedini Ponomarenko p. 82 |
| 1990 | Ommatinae Hong and Wang p. 105 |
| 1995 | Ommatinae Lawrence and Newton, Jr. |
| 2000 | Tetraphalerus brevicapitis Ponomarenko and Martínez-Delclòs p. 48 figs. 1.3, 2.1 |
| 2006 | Ommatinae Soriano and Delclos |
| 2009 | Notocupedinae Tan and Ren p. 74 |
| 2009 | Ommatinae Tan and Ren p. 91 |
| 2009 | Tetraphalerini Tan and Ren p. 112 |
| 2011 | Lithocupedini Bouchard et al. p. 96 |
| 2011 | Ommatinae Bouchard et al. p. 96 |
| 2012 | Lithocupedini Tan et al. |
| 2012 | Pronotocupedini Tan et al. |
| 2012 | Tetraphalerini Tan et al. |
| 2012 | Pronotocupedini Tan et al. p. 16 |
| 2013 | Ommatinae Jarzembowski et al. p. 139 |
| 2014 | Ommatinae Ponomarenko p. 1658 |
| 2016 | Ommatinae Jarzembowski et al. p. 114 |
| 2017 | Ommatinae Cai and Huang p. 279 |
| 2017 | Ommatinae Jarzembowski et al. p. 85 |
| 2017 | Tetraphalerini Jarzembowski et al. p. 85 |
| 2017 | Ommatinae Yamamoto p. 35 |
| 2019 | Ommatinae Jarzembowski et al. p. 316 |
| 2019 | Ommatinae Tihelka et al. p. 571 |
| 2020 | Ommatinae Kirejtshuk p. 22 |
| 2022 | Ommatinae Cai et al. |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Subfm. Ommatinae Sharp and Muir 1912
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G. †Allophalerus Kirejtshuk 2020
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†Allophalerus antiquus Ponomarenko 1964
†Allophalerus aphaleratus Ponomarenko 1969
†Allophalerus bontsaganensis Ponomarenko 1997
†Allophalerus incertus Ponomarenko 1969
†Allophalerus latus Tan et al. 2007
†Allophalerus maximus Ponomarenko 1968
†Allophalerus okhotensis Ponomarenko 1993
†Allophalerus tenuipes Ponomarenko 1964
†Allophalerus verrucosus Ponomarenko 1966
G. †Blapsium Westwood 1854
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†Blapsium egertoni Westwood 1854
Invalid names: Blapsidium Phillips 1871 [synonym]
G. †Brochocoleus Hong 1982
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†Brochocoleus alatus Ponomarenko 1994
†Brochocoleus cossyphus Ponomarenko 1994
†Brochocoleus keenani Jarzembowski et al. 2013
†Brochocoleus maculatus Whalley 1985
†Brochocoleus maximus Jarzembowski et al. 2013
†Brochocoleus planus Ponomarenko 1994
†Brochocoleus punctatus Hong 1982
†Brochocoleus rostratus Ponomarenko 1999
†Brochocoleus sacheonensis Lee et al. 2023
†Brochocoleus tobini Jarzembowski et al. 2013
G. †Burmocoleus Kirejtshuk 2020
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†Burmocoleus prisnyi Kirejtshuk 2020
†Burmocoleus zhiyuani Liu et al. 2017
G. †Cionocoleus Ren 1995
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†Cionocoleus cervicalis Tan et al. 2007
†Cionocoleus elizabethae Jarzembowski et al. 2013
†Cionocoleus jepsoni Jarzembowski et al. 2013
†Cionocoleus longicapitis Soriano and Delclos 2006
†Cionocoleus magicus Ren 1995
†Cionocoleus minimus Jarzembowski et al. 2013
†Cionocoleus olympicus Jarzembowski et al. 2013
†Cionocoleus ommamimus Ponomarenko 1997
†Cionocoleus planiusculus Tan et al. 2007
†Cionocoleus punctatus Martynov 1926
†Cionocoleus sibiricus Ponomarenko 2000
†Cionocoleus tanae Jarzembowski et al. 2013
†Cionocoleus watsoni Jarzembowski et al. 2013
Tr. †Clessidrommatini Jarzembowski et al. 2017
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G. †Lepidomma Jarzembowski et al. 2019
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†Lepidomma beuteli Song et al. 2022
†Lepidomma jarzembowskii Li and Cai 2020
†Lepidomma longisquama Li and Cai 2020
†Lepidomma tianae Jarzembowski et al. 2019
G. †Diluticupes Ren 1995
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†Diluticupes applanatus Tan and Ren 2009
†Diluticupes crowsonae Jarzembowski et al. 2013
†Diluticupes impressus Ren 1995
†Diluticupes magnus Tan and Ren 2009
†Diluticupes minor Ponomarenko 2000
†Diluticupes validus Tan and Ren 2009
†Diluticupes yangshuwanziensis Jarzembowski et al. 2013
G. †Echinocups Kirejtshuk and Jarzembowski 2020
G. †Lithocupes Ponomarenko 1966
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†Lithocupes gigas Ponomarenko 1969
†Lithocupes incertus Ponomarenko 1966
†Lithocupes punctatus Ponomarenko 1969
G. †Monticupes Ren 1995
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†Monticupes curtinervis Tan et al. 2007
†Monticupes decorosus Tan et al. 2012
†Monticupes fentaiensis Ren 1995
†Monticupes surrectus Ren 1995
G. †Notocupoides Ponomarenko 1966
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†Notocupoides capitatus Ponomarenko 1966
†Notocupoides fasciatus Ponomarenko 1966
†Notocupoides triassicus Ponomarenko 1966
G. †Odontomma Ren et al. 2006
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†Odontomma patula Ponomarenko 1985
†Odontomma sulcatum Tan et al. 2007
†Odontomma trachylaenum Ren et al. 2006
Tr. Ommatini Sharp and Muir 1912
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G. Omma Newman 1839
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†Omma aberratum Ponomarenko 1968
†Omma altajense Ponomarenko 1997
†Omma ancistrodontum Tan et al. 2012
†Omma antennatum Ponomarenko 1997
†Omma avus Ponomarenko 1969
†Omma brevipes Deichmüller 1886
†Omma davidbatteni Jarzembowski et al. 2020
†Omma daxishanense Cai and Huang 2017
†Omma delicatum Tan et al. 2012
†Omma elongatum Brodie 1845
†Omma forte Li and Cai 2021
†Omma gobiense Ponomarenko 1997
†Omma grande Ponomarenko 1964
†Omma janetae Kirejtshuk 2020
†Omma jurassicum Ponomarenko 1968
†Omma liassicum Crowson 1962
†Omma lii Jarzembowski et al. 2016
†Omma longicolle Ponomarenko 1997
†Omma manukyani Kirejtshuk 2020
†Omma pilosum Ponomarenko 1964
†Omma sibiricum Ponomarenko 1966
†Omma zitteli Oppenheim 1888
Invalid names: Chalepocarabus Handlirsch 1906 [synonym], Cionocups Kirejtshuk 2020 [synonym], Ommomima Ponomarenko 1964 [synonym], Procarabus Oppenheim 1888 [synonym], Pyrochroophana Handlirsch 1906 [synonym]
G. †Paraodontomma Yamamoto 2017
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†Paraodontomma burmitica Yamamoto 2017
†Paraodontomma leptocristatum Li et al. 2021
†Paraodontomma szwedoi Jarzembowski et al. 2018
G. †Pareuryomma Tan et al. 2012
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†Pareuryomma angustum Tan et al. 2007
†Pareuryomma cardiobasis Tan et al. 2012
†Pareuryomma tylodes Tan et al. 2006
Invalid names: Euryomma Tan et al. 2006 [replaced]
G. †Polyakius Kirejtshuk 2020
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†Polyakius alberti Kirejtshuk 2020
†Polyakius pubescens Kirejtshuk 2020
G. †Stegocoleus Jarzembowski and Wang 2016
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†Stegocoleus arkonus Tihelka et al. 2019
†Stegocoleus caii Jarzembowski and Wang 2016
†Stegocoleus lawrencei Tihelka et al. 2019
G. Tetraphalerus Waterhouse 1901
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†Tetraphalerus brevis Ponomarenko 1964
†Tetraphalerus collaris Ponomarenko 1997
†Tetraphalerus glabratus Ponomarenko 1997
†Tetraphalerus largicoxa Lin 1986
†Tetraphalerus mongolicus Ponomarenko 1986
†Tetraphalerus notatus Ponomarenko 1997
†Tetraphalerus penalveri Soriano and Delclos 2006
†Tetraphalerus ponomarenkoi Soriano and Delclos 2006
Invalid names: Lithocupedini Ponomarenko 1969 [invalid subgroup], Notocupedinae Ponomarenko 1966 [synonym], Pronotocupedini Tan et al. 2012 [invalid subgroup], Tetraphalerini Crowson 1962 [invalid subgroup], Tetraphalerus brevicapitis Ponomarenko and Martínez-Delclòs 2000 [nomen dubium]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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| Source: c = class, p = phylum | |||||
| References: Bush and Bambach 2015, Kiessling 2004 | |||||
Age range
Maximum range based only on fossils: base of the Ladinian to the top of the Priabonian or 241.46400 to 33.90000 Ma
Minimum age of oldest fossil (stem group age): 237.0 Ma
Minimum age of oldest fossil (stem group age): 237.0 Ma
Collections (91 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Ladinian | Kyrgyzstan (Osh) | Lithocupes gigas (106088) Lithocupes incertus (110767) Lithocupes punctatus (110763) Notocupoides fasciatus, Notocupoides capitatus (106090) Notocupoides triassicus (113468) | |
| Rhaetian | United Kingdom | Omma liassica (133036) | |
| Rhaetian | United Kingdom (England) | Omma liassica (113957 114677) | |
| Rhaetian - Early/Lower Hettangian | United Kingdom (England) | Omma liassica (124688) | |
| Planorbis | United Kingdom (England) | Omma liassica (123982) | |
| Hettangian - Sinemurian | Kyrgyzstan (Tonskiy) | Omma avus (105087) Tetraphalerus incertus (113432) | |
| Obtusum | United Kingdom (England) | Liassocupes maculatus (114638) Liassocupes parvus (114639) Omma liassica (114637) | |
| Pliensbachian | Kyrgyzstan (Osh) | Eurydictyon conspicuum, Tetraphalerus aphaleratus (122971) | |
| Toarcian | Australia (Western Australia) | Tetraphalerus sp. (114239) | |
| Toarcian - Aalenian | China (Hunan) | Tetraphalerus largicoxa (114577) | |
| Bajocian - Bathonian | Mongolia (Bayan-Khongor) | Tetraphalerus glabratus (124339) Tetraphalerus glabratus, Omma altajense (124340) | |
| Bathonian | United Kingdom (Oxfordshire) | Blapsium egertoni (153647) | |
| Callovian | Russian Federation (Tyumenskaya) | Notocupes patulus (125470) | |
| Callovian - Oxfordian | Kazakhstan (Chimkent) | Carabopteron punctatum (115792) Omma jurassicum (105970 106101) Ommomima pilosa (112846) Ommomima pilosa, Tetraphalerus brevis (105971) Tetraphalerus antiquus, Tetraphalerus tenuipes (113402) Tetraphalerus grandis (31311) Tetraphalerus maximus (115717) Tetraphalerus maximus, Omma aberratum (106102) | |
| Callovian - Oxfordian | China (Nei Mongol) | Brochocoleus magnus, Brochocoleus validus, Brochocoleus applanatus (113371) Pareuryomma ancistrodonta, Omma delicata, Tetraphalerus decorosus (113366) | |
| Oxfordian | China (Liaoning) | Omma daxishanense (185124) | |
| Early/Lower Tithonian | Germany (Bayern) | Omma brevipes (123144) Pyrochroa brevipes (133578) Pyrochroophana maior, Buprestites suprajurensis, Elaterites robustus, Procarabus zitteli (123985) | |
| Early/Lower Tithonian | Germany (Bavaria) | Pyrochroophana brevipes (122586 139876) | |
| Tithonian | Australia (New South Wales) | Rhopalomma stefaniae (128121) | |
| Tithonian | Mongolia (Dund-Gov’ Aymag) | Omma gobiense (122321) | |
| Berriasian | United Kingdom (England) | Carabus elongatus (105539) | |
| Late/Upper Berriasian | United Kingdom (England) | Brochocoleus maximus, Cionocoleus jepsoni (160681) Brochocoleus sp. (122909) | |
| Late/Upper Hauterivian | United Kingdom (England) | Chalepocarabus elongatus, Brochocoleus tobini (119697) Cionocoleus minimus (106417) | |
| Late/Upper Hauterivian | United Kingdom (Surrey) | Cionocoleus minimus (74972) | |
| Hauterivian - Barremian | Mongolia (Bayankhongor) | Tetraphalerus bontsaganensis, Tetraphalerus notatus (113357) | |
| Hauterivian - Barremian | Mongolia | Brochocoleus planus (125191) | |
| Barremian | Mongolia | Tetraphalerus longicollis, Brochocoleus alatus (124343) | |
| Early/Lower Barremian | Spain | Cionocoleus longicapitis (123437) Tetraphalerus brevicapitis (152691) Tetraphalerus penalveri (152690) | |
| Early/Lower Barremian | United Kingdom (England) | Brochocoleus crowsonae, Brochocoleus keenani, Cionocoleus elizabethae, Cionocoleus watsoni (45193) | |
| Late/Upper Barremian | China (Liaoning) | Cionocoleus cervicalis (140491) Cionocoleus planiusculus (152003) Euryomma tylodes (111717) Odontomma trachylaena (152052) Pareuryomma cardiobasis (141615) Tetraphalerus latus, Tetraphalerus curtinervis, Brochocoleus sulcatus, Brochocoleus angustus (111710) | |
| Late/Upper Barremian | China (Nei Mongol) | Cionocoleus magicus, Brochocoleus impressus, Brochocoleus sulcatus, Brochocoleus yangshuwanziensis, Cionocoleus cervicalis, Cionocoleus tanae, Cionocoleus olympicus (103654) | |
| Late/Upper Barremian | Spain (Castilla-La Mancha) | Tetraphalerus ponomarenkoi (28006) | |
| Early/Lower Aptian | Russian Federation | Brochocoleus minor, Cionocoleus sibiricus (123220) | |
| Aptian | Russian Federation (Buryatia) | Omma sibiricum, Tetraphalerus verrucosus (105536) | |
| Aptian | Mongolia | Brochocoleus cossyphus (125186) Cionocoleus ommamimus (105796 124344) Tetraphalerus bontsaganensis (124341) Tetraphalerus bontsaganensis, Brochocoleus alatus (124342) Tetraphalerus bontsaganensis, Tetraphalerus longicollis, Brochocoleus cossyphus, Brochocoleus alatus, Brochocoleus planus (113359) Tetraphalerus bontsaganensis, Tetraphalerus longicollis, Tetraphalerus notatus, Omma antennatum, Brochocoleus cossyphus, Brochocoleus alatus, Brochocoleus planus (113355) Tetraphalerus collaris, Tetraphalerus bontsaganensis, Tetraphalerus longicollis, Tetraphalerus notatus, Brochocoleus cossyphus, Brochocoleus planus (113356) Tetraphalerus longicollis (103273) | |
| Aptian | China (Gansu) | Brochocoleus punctatus (155034) | |
| Aptian | Mongolia (Govi-Altai) | Tetraphalerus mongolicus (122332) | |
| Late/Upper Aptian | China (Beijing Shi) | Cionocoleus magicus, Monticupes surrectus, Monticupes fentaiensis, Diluticupes impressus (106965) | |
| Albian | South Korea | Brochocoleus punctatus (184781) Omma sp., Brochocoleus sacheonensis (232407) | |
| Early/Lower Cenomanian | Myanmar | Brochocoleus zhiyuani (165561) Burmocoleus prisnyi, Omma janetae, Polyakius alberti (207883) Cionocups manukyani (172900) Lepidomma beuteli (224419) Omma lii (183293) Paraodontomma burmitica (111688) Polyakius pubescens (192152) Stegocoleus caii (179937 179938) Stegocoleus caii, Omma lii, Brochocoleus caseyi, Paraodontomma szwedoi, Lepidomma tianae, Stegocoleus arkonus, Stegocoleus lawrencei, Burmocoleus sp., Burmocoleus prisnyi, Omma davidbatteni, Paraodontomma leptocristatum, Omma forte, Omma manukyani, Lepidomma jarzembowskii, Lepidomma longisquama, Brochocoleus sp. A, Lepidomma tianae (161726) | |
| Turonian | Kazakhstan | Brochocoleus rostratus (103274) | |
| Late/Upper Coniacian | Russian Federation (Khabarovsk) | Tetraphalerus okhotensis (123528) | |
| Priabonian | United Kingdom (England) | Tetraphalerites oligocenicus (123959) |