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Leucosiidae

Malacostraca - Decapoda - Leucosiidae

Taxonomy
Leucosiidae was named by Samouelle (1819). It is extant. Its type is Leucosia.

It was assigned to Oxystomata by Rathbun (1918), Rathbun (1926); to Calappoidea by Tiwari et al. (1997); and to Leucosioidea by Karasawa (1993), Collins et al. (2003), Todd and Collins (2005), Schweitzer et al. (2010), Luque et al. (2017), Karasawa et al. (2019).

Synonymy list
YearName and author
1819Leucosiidae Samouelle
1918Leucosiidae Rathbun p. 161
1926Leucosiidae Rathbun p. 75
1993Leucosiidae Karasawa p. 75
1997Leucosiidae Tiwari et al. p. 128
2003Leucosiidae Collins et al. p. 202
2005Leucosiidae Todd and Collins p. 71
2010Leucosiidae Schweitzer et al. p. 87
2017Leucosiidae Luque et al. p. 12
2019Leucosiidae Karasawa et al. p. 1

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaLatreille 1829
—Deuteropoda
—Mandibulata
—PancrustaceaZrzavý and Štys 1997
—Hymenostraca(Rolfe 1969)
superclassMulticrustaceaRegier et al. 2010
classMalacostraca()
RankNameAuthor
subclassEumalacostracaGrobben 1892
—CaridoidaHessler 1982
orderDecapoda
suborderPleocyemataBurkenroad 1963
—Reptantia(Boas)
—MeiuraSaint Laurent 1980
infraorderBrachyura()
—Eubrachyurade Saint Laurent 1980
—HeterotremataGuinot 1977
superfamilyLeucosioideaSamouelle 1819
familyLeucosiidaeSamouelle 1819
familyLeucosiidaeSamouelle 1819

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. Leucosiidae Samouelle 1819
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Subfm. Cryptocneminae Stimpson 1907
+
G. Cryptocnemus Stimpson 1858
+
Subfm. Ebaliinae Stimpson 1871
+
G. Arcania Leach 1817
+
Arcania cornuta MacGilchrist 1905
Arcania elongata Yokoya 1933
Arcania globata Stimpson 1858
Arcania gracilis Henderson 1893
Arcania septemspinosa Fabricius 1787
†Arcania shuishuni Hu and Tao 1985
Arcania siamensis Rathbun 1909
G. †Cabrillia Nyborg et al. 2018
+
†Cabrillia hendyi Nyborg et al. 2018
G. Ebalia Leach 1817
+
Ebalia cranchii Leach 1817
Ebalia crassipes Bell 1855
Ebalia cryptocnemoides Takeda and Miyake 1972
Ebalia dentifrons Miers 1886
†Ebalia deshayesi Lucas 1846
†Ebalia ferganensis Ilyin and Alekseev 2005
†Ebalia fucinii Ristori 1893
†Ebalia globulosa Müller 1976
†Ebalia hungarica Müller 1974
Ebalia intermedia Miers 1886
†Ebalia jacqueshermani van Bakel et al. 2003
†Ebalia kakegawaensis Karasawa 1993
†Ebalia lamarmorai Lorenthey 1909
Ebalia longimana Ortmann 1892
†Ebalia meulenkampi Muller and Georgiades 1984
†Ebalia multiangulata Muller 1993
†Ebalia noetlingi Karasawa 2008
+
Invalid names: Ebalia tuberculata Noetling 1901 [replaced]
Ebalia nux Milne-Edwards 1883
†Ebalia oersi Müller 1978
Ebalia spinifera Miers 1886
†Ebalia spinosa Tiwari et al. 1997
Ebalia tuberculosa Milne-Edwards 1873
Ebalia tuberosa Pennant 1777 [Pennant's nut crab]
Ebalia tuberosus Pennat 1777
Ebalia tumefacta Montagu 1808 [Bryer's nut crab]
†Ebalia vahldieki Forster and Mundlos 1992
†Ebalia vanstraeleni Bachmayer 1953
G. Heteronucia Alcock 1896
+
G. Hiplyra Galil 2009
+
Hiplyra platycheir De Haan 1841
G. Ilia Leach 1817
+
Ilia nucleus Linnaeus 1758
†Ilia ornata Randall 1839
G. Iliacantha Stimpson 1871
+
Iliacantha globosa Stimpson 1981
†Iliacantha liodactylus Rathbun 1938
†Iliacantha panamanica Todd and Collins 2005
G. Ixa Leach 1816
+
Ixa cylindrus Fabricius 1777
+
†Ixoides miocenicus Garassino et al. 2024
G. Leucosilia Bell 1855
+
†Leucosilia bananensis Rathbun 1918
Leucosilia jurinei Saussure 1853
Leucosilia jurinii De Saussure 1853
G. †Leucosiraja De Angeli et al. 2017
+
†Leucosiraja manta De Angeli et al. 2017
G. Lyphira Galil 2009
+
Lyphira heterograna Ortmann 1892
G. Merocryptus Milne-Edwards 1873
+
†Merocryptus altavillensis De Angeli et al. 2010
Merocryptus lambriformis Poore and Ahyong 2004
G. Myra Leach 1817
+
Myra affinis Bell 1855
†Myra amoena Bohm 1922
†Myra arachnoides Bohm 1922
†Myra brevisulcata Morris and Collins 1991
Myra celeris Galil 2001
†Myra emarginata Glaessner 1928
†Myra fijiensis Rathbun 1934
Myra fugax Fabricius 1798 [fleeting purse crab]
Myra mammillaris Bell 1855
†Myra paenitensis Via 1941
†Myra subcarinata Morris and Collins 1991
†Myra trispinosa Morris and Collins 1991
†Myra wenjii Hu and Tao 1985
G. Myropsis Stimpson 1871
G. Nucia Dana 1852
+
†Nucia baripadensis Bachmayer and Mohanti 1973
†Nucia borneoensis Morris and Collins 1991
†Nucia calculoides Morris and Collins 1991
†Nucia coxi Morris and Collins 1991
†Nucia fennemai Bohm 1922
†Nucia platyspinosa Collins et al. 2003
Nucia speciosa Dana 1852
G. Nursia Leach 1817
+
†Nursia bilobata Karasawa et al. 2008
Nursia japonica Sakai 1935
G. Paranursia Serene and Soh 1976
+
†Paranursia acharyai Bachmayer and Mohanti 1973
G. Persephona Leach 1817
+
†Persephona bigranulata Rathbun 1926
†Persephona enigmatica Todd and Collins 2005
†Persephona invalida Rathbun 1929
†Persephona manningi Todd and Collins 2005
†Persephona niemeyeri Blow 2003
†Persephona prepunctata Rathbun 1919
Persephona punctata Linnaeus 1758
Persephona punctatus Linnaeus 1758
+
Invalid names: Persephona latreillii Leach 1817 [synonym]
†Persephona rodesae Blow 2003
Invalid names: Guaia Leach 1817 [synonym]
G. Philyra Leach 1817
+
†Philyra alveola Hu and Tao 2000
†Philyra collinsi Karasawa 2010
†Philyra cranium Desmarest 1822
†Philyra ferrica Hu and Tao 1996
†Philyra granulosa Morris and Collins 1991
†Philyra hayasakai Karasawa and Inoue 1992
†Philyra hormozganensis Vega et al. 2010
†Philyra miyamotoi Karasawa and Kishimoto 1996
†Philyra nishimotoi Karasawa 1989
Philyra olivacea Rathbun 1909
†Philyra plana Karasawa 1989
Philyra scabriuscula Fabricius 1793
†Philyra shihcheni Hu and Tao 1979
Philyra syndactyla Ortmann 1892
†Philyra tanakai Karasawa 1993
†Philyra tridentata Karasawa 1993
†Philyra trusanensis Collins et al. 2003
G. Pseudophilyra Miers
+
†Pseudophilyra elongatella Hu and Tao 1996
G. Pyrhila Galil 2009
+
Pyrhila pisum De Haan 1841
G. Randallia Stimpson 1857
+
†Randallia jingomao Hu and Tao 2004
†Randallia ornata Randall 1839
†Randallia pleistocenica Rathbun 1926
†Randallia prolanata Hu and Tao 1996
†Randallia saitoensis Karasawa 1993
†Randallia strouhali Bachmayer 1953
G. Ryphila Galil 2009
G. Speloeophorus Milne Edwards 1865
+
†Speloeophorus subcircularis Todd and Collins 2005
G. Tokoyo Galil 2003
+
Tokoyo eburnea Alcock 1896
†Tokoyo trinucloidea Hu and Tao 1996
G. Uhlias Stimpson 1871
+
Uhlias limbatus Stimpson 1871
Subfm. Leucosiinae Samouelle 1819
+
G. Euclosiana Galil and Ng 2010
+
†Euclosiana obtusifrons de Haan 1841
Invalid names: Euclosia Galil 2003 [replaced]
G. Leucosia Weber 1795
+
Leucosia anatum Herbst 1783
†Leucosia calcarata Collins et al. 2003
Leucosia craniolaris Linnaeus 1758
†Leucosia longiangulata Morris and Collins 1991
†Leucosia martini Karasawa 2008
†Leucosia ovalata Hu and Tao 1996
Leucosia perlata De Haan 1841
†Leucosia persica Vega et al. 2010
Leucosia pubescens Miers 1877
†Leucosia serenei Morris and Collins 1991
†Leucosia subrhomboidalis Desmarest 1822
†Leucosia subrhomboidea Hu and Tao 1996
†Leucosia taiwanica Hu and Tao 1996
†Leucosia takamii Karasawa 1993
†Leucosia tricarinata Martin 1879
†Leucosia tutongensis Morris and Collins 1991
G. †Ristoria Garassino 2012
+
†Ristoria pliocaenica Ristori 1891
G. Seulocia Galil 2005
G. Urnalana Galil 2005
+
Urnalana haematosticta Adams and White 1848
Subfm. Philyrinae Leach 1817
+
G. Bellidilia Kinahan 1856
+
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Locomotion: actively mobilec
Life habit: semi-infaunalsuperf
Diet: carnivoreo
Vision: well-developedc
Created: 2022-11-16 16:36:32
Modified: 2022-11-16 16:36:32
Source: superf = superfamily, o = order, c = class
References: Turnsek 1997, Aberhan 1992, Fearon and Clapham 2023

Age range

Maximum range based only on fossils: base of the Ypresian to the top of the Holocene or 56.00000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 47.8 Ma

Collections (159 total)


Time interval Ma Country or state Original ID and collection number
Ypresian56.0 - 47.8Italy Leucosiidae indet. (207470)
Priabonian38.0 - 33.9Italy Leucosiidae indet. (207550)
Rupelian33.9 - 28.1Italy (Vicenza) Merocryptus altavillensis (207617)
Oligocene33.9 - 23.03USA (Oregon) Persephona bigranulata (139538)
Late/Upper Oligocene - Early/Lower Miocene28.4 - 15.97Cuba (Matanzas) Iliacantha liodactylus (228535)
Aquitanian23.03 - 20.44Mexico (Chiapas) Iliacantha panamica (133885)
Early/Lower Miocene23.03 - 15.97Brazil (Para) Randallia sp. (65162)
Early/Lower Miocene23.03 - 15.97India Ebalia spinosa, Ebalia tuberculata (78641)
Early/Lower Miocene23.03 - 15.97India (Mizoram) Ebalia spinosa, Ebalia tuberculata (42760)
Early/Lower Miocene23.03 - 15.97Cuba (Sancti Spiritus) Persephona sp. (228530)
Early/Lower Miocene23.03 - 15.97Malaysia (Sabah) Nucia coxi (173566 226228)
Miocene23.03 - 5.333Italy Philyra hayasakai (147814)
Miocene23.03 - 5.333Indonesia Leucosia tricarinata, Leucosia unidentata, Leucosia obtusifrons (81717)
Miocene23.03 - 5.333Japan (Shindon) Philyra miyamotoi (127687)
Miocene23.03 - 5.333Sri Lanka Leucosia unidentata (13449)
Miocene23.03 - 5.333Japan Philyra hayasakai (147808 147809 147810 147811 147812 147817 147818 147819 147820 147821)
Miocene - Late/Upper Pliocene23.03 - 2.588USA (North Carolina) Persephona punctata (144073)
Neogene23.03 - 2.58Fiji Myra fijiensis (136017)
Burdigalian20.44 - 15.97Malaysia (Sarawak) Myra trispinosa, Leucosia calcarata, Leucosia longiangulata (175184)
Burdigalian20.44 - 15.97Venezuela (Falcon) Iliacantha sp. (228724)
Langhian15.97 - 13.82Japan (Okayama) Philyra plana (127702)
Langhian - Serravallian15.97 - 11.62Iran (Hormozgan) Arcania sp., Leucosia persica, Philyra hormozganensis (101700)
Serravallian13.82 - 11.62Panama (Colon) Leucosilia bananensis (228718)
Serravallian13.82 - 11.62Trinidad and Tobago Persephona punctata (171483)
Serravallian - Tortonian13.82 - 7.246Malaysia (Sarawak) Nucia borneoensis, Nucia platyspinosa, Philyra trusanensis, Philyra granulosa (175181)
Badenian13.65 - 12.7Hungary Ebalia hungarica (138812 140154 149366 207750)
Tortonian11.62 - 7.246Dominican Republic Persephona prepunctata (58817)
Tortonian11.62 - 7.246Panama Leucosilia bananensis (145520)
Tortonian11.62 - 7.246Iran Ixoides miocenicus (234649)
Tortonian11.62 - 7.246Malaysia (Sabah) Myra subcarinata (185853 185854) Myra subcarinata, Nucia platyspinosa (185833 185834 185849) Myra subcarinata, Nucia platyspinosa, Leucosia serenei, Philyra granulosa (185835) Nucia platyspinosa (185846 185847 185850 185851)
Tortonian - Messinian11.62 - 5.333Philippines (Batangas) Philyra acutidens, Pseudophilyra granulimarginata, Nursia bilobata (224585)
Tortonian - Messinian11.62 - 5.333Panama (Bocas del Toro) Iliacantha panamanica, Leucosilia bananensis (228707)
Late/Upper Miocene11.608 - 5.333Venezuela (Falcon) Iliacantha sp. (228732)
Late/Upper Miocene11.608 - 5.333USA (South Carolina) Persephona punctata (144074)
Late/Upper Miocene11.608 - 5.333Dominican Republic Persephona prepunctata (228607 228608 228609 228610 228611 228612 228613 228616 228620 228630 228631 228632 228633 228634 228635 228636)
Late/Upper Miocene11.608 - 5.333Brunei Darussalam Myra subcarinata, Myra trispinosa, Leucosia serenei, Leucosia tutongensis (173581) Myra subcarinata, Myra trispinosa, Nucia borneoensis, Leucosia serenei, Leucosia tutongensis, Philyra granulosa (173589) Myra trispinosa, Nucia calculoides, Leucosia tutongensis (173586) Nucia borneoensis (173591) Philyra granulosa (173574 173590)
Late/Upper Miocene11.608 - 5.333Indonesia Leucosia tricarinata, Leucosia obtusifrons, Myra fugax (70098) Leucosia tricarinata, Myra fugax (70099)
Late/Upper Miocene11.608 - 5.333Malaysia Myra brevisulcata, Nucia calculoides, Leucosia longiangulata (226226) Nucia borneoensis (226227)
Late/Upper Miocene11.608 - 5.333Mexico (Veracruz) Persephona sp. (228595)
Late/Upper Miocene - Early/Lower Pliocene11.608 - 3.6Dominican Republic Iliacantha sp. (228615) Persephona prepunctata (228622 228623)
Late/Upper Miocene - Early/Lower Pliocene11.608 - 3.6USA (Virginia) Persephona punctata (144067 144069 144070 144071)
Late/Upper Miocene - Pliocene11.608 - 2.588Colombia (Atlántico) Persephona enigmatica (228493)
Messinian7.246 - 5.333Greece (Crete) Ebalia meulenkampi (148521)
Early/Lower Pliocene5.333 - 3.6Panama (Bocas del Toro) Leucosilia bananensis (228683)
Early/Lower Pliocene5.333 - 3.6Japan (Miyazaki) Randallia saitoensis (127696)
Zanclean5.333 - 3.6Panama (Bocas del Toro) Leucosilia bananensis (228688) Persephona enigmatica (228692)
Zanclean5.333 - 3.6Netherlands Antilles (Curacao) Myropsis quinquespinosa (228642)
Pliocene5.333 - 2.588Italy (Biella) Ilia pliocaenica (127497)
Pliocene5.333 - 2.588USA (California) Persephona invalida (135776)
Pliocene5.333 - 2.588Italy Ilia pliocaenica (221587)
Pliocene5.333 - 2.588USA (North Carolina) Persephona punctata (144072)
Pliocene5.333 - 2.588Panama (Bocas del Toro) Leucosilia bananensis (228694)
Pliocene5.333 - 2.588Italy (Siena) Ebalia deshayesi (221595) Ilia pliocenica (221592 221593)
Zanclean - Middle Pleistocene5.333 - 0.129Italy (Piacenza) Ebalia cranchii, Ilia pliocaenica (127162)
Pliocene - Pleistocene5.333 - 0.0117Italy (Livorno) Ebalia cranchii, Ebalia fucinii (221590)
Late/Upper Pliocene3.6 - 2.588Japan (Okinawa) Myra fugax (42383)
Late/Upper Pliocene3.6 - 2.588Jamaica Persephona punctata (228744)
Late/Upper Pliocene3.6 - 2.588Costa Rica (Limon) Leucosilia bananensis (228540 228542) Persephona enigmatica (228538)
Late/Upper Pliocene3.6 - 2.588Japan (Shizuoka) Ebalia kakegawaensis (202454) Leucosia sp., Philyra tanakai, Philyra platychira (127710) Leucosia sp., Philyra tridentata, Leucosia takamii, Philyra platychira (202449) Leucosia takamii (127713) Philyra tanakai, Philyra platychira (202451)
Piacenzian3.6 - 2.588Costa Rica Leucosilia bananensis (131313 131340 226680)
Piacenzian3.6 - 2.588Belgium Ebalia cranchii, Ebalia jacqueshermani (221313) Ebalia tumefacta (221312)
Piacenzian3.6 - 2.588Panama (Bocas del Toro) Iliacantha sp. (228693) Persephona manningi (228703) Speloeophorus subcircularis (228695)
Late/Upper Pliocene - Early/Lower Pleistocene3.6 - 0.781Costa Rica (Limon) Persephona sp. (228545)
Early/Lower Pleistocene2.588 - 0.781Costa Rica (Puntarenas) Speloeophorus subcircularis (228536)
Gelasian - Calabrian2.588 - 0.774Philippines (Mindanao) Leucosia martini (224586)
Pleistocene2.588 - 0.0117Japan (Okinawa-jima) Myra fugax (127689)
Pleistocene2.588 - 0.0117USA (North Carolina) Persephona punctata (159988)
Pleistocene2.588 - 0.0117USA (South Carolina) Persephona punctata (150656)
Pleistocene2.588 - 0.0117USA (California) Randallia ornata (138720) Randallia ornata, Randallia pleistocenica (138719)
Pleistocene - Holocene2.588 - 0.0USA (California) Randallia ornata (138721 138722 138821) Randallia pleistocenica (138724)
Calabrian1.8 - 0.774Italy Ebalia nux (234392 234395 234396 234397 234398 234399 234400 234401 234402 234403) Ebalia sp., Ebalia nux (234394)
Middle Pleistocene0.774 - 0.129USA (California) Randallia ornata (138723)
Middle Pleistocene0.774 - 0.129Japan (Aichi) Leucosia haematosticta (127686)
Middle Pleistocene0.774 - 0.129Japan (Ishikawa) Leucosia sp. (202458)
Late/Upper Pleistocene0.129 - 0.0117USA (California) Randallia ornata (59063)
Late/Upper Pleistocene0.129 - 0.0117Panama Leucosiidae indet., Leucosilia jurinei (134957)
Late/Upper Pleistocene0.129 - 0.0117Jamaica Persephona sp. (185031) Uhlias limbatus (228847)
Tarantian0.129 - 0.0117Jamaica Persephona punctata (224549)
Holocene0.0117 - 0.0USA (South Carolina) Persephona punctata (146572)
Holocene0.0117 - 0.0USA (Florida) Persephona punctata (146566)
Holocene0.0117 - 0.0Panama Persephona edwardsii, Leucosilia jurinii (146569)