PBDB Taxon

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Eulipotyphla

Mammalia

Taxonomy

Eulipotyphla is an unranked clade. It is extant.

It was considered an invalid subgroup of Lipotyphla by Asher et al. (2005), Asher and Helgen (2010).

It was assigned to Laurasiatheria by Waddell et al. (2001), Madsen et al. (2001), Douady et al. (2002), Silcox et al. (2010).

Subtaxa

Aethomylos, Erinaceidae (syn. Acanthionidae), Erinaceoidae, Erinaceomorpha, Ghamidtherium, Pantolestoidae, Solenodontidae, Soricomorpha

Sister taxa

Erinaceomorpha

Synonyms

Lipotyphla was named by Haeckel (1866). It is extant.

It was reranked as the suborder Lipotyphla by Gregory (1910), Jaekel (1911), Hay (1930); it was synonymized subjectively with Eulipotyphla by Asher and Helgen (2010).

It was assigned to Insectivori by Jaekel (1911); to Insectivora by Gregory (1910), Hay (1930); to Eutheria by Hooker (1986); to Mammalia by Tabuce et al. (2005); to Insectivora by Kielan-Jaworowska et al. (2004), Lopatin (2006), Scott (2006); to Placentalia by Wible et al. (2007); and to Eutheria by Vianey-Liaud et al. (2014).

Synonymy list

Year	Name and author
1866	Lipotyphla Haeckel
1910	Lipotyphla Gregory p. 464
1911	Lipotyphla Jaekel p. 210
1930	Lipotyphla Hay p. 424
1986	Lipotyphla Hooker p. 223
2001	Eulipotyphla Madsen et al. p. 610
2001	Eulipotyphla Waddell et al. p. 148
2002	Eulipotyphla Douady et al. p. 204
2004	Lipotyphla Kielan-Jaworowska et al. pp. 494, 506
2005	Lipotyphla Tabuce et al. p. 390
2006	Lipotyphla Lopatin
2006	Lipotyphla Scott p. 1697
2007	Lipotyphla Wible et al.
2010	Eulipotyphla Silcox et al.
2014	Lipotyphla Vianey-Liaud et al. p. 576

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota

Rank	Name	Author
subclass	Synapsida
—	Therapsida	()
infraorder	Cynodontia	()
—	Mammaliamorpha
—	Mammaliaformes
class	Mammalia
—	Cladotheria
—	Zatheria
subclass	Tribosphenida	()
subclass	Theria
—	Eutheria	()
—	Placentalia
—	Boreoeutheria
—	Laurasiatheria
—	Eulipotyphla
—	Eulipotyphla

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. Eulipotyphla

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Unr. Erinaceomorpha Gregory 1910

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Fm. †Chambelestidae Gheerbrant and Hartenberger 1999

G. †Chambilestes Gheerbrant and Hartenberger 1999

†Chambilestes foussanensis Gheerbrant and Hartenberger 1999

G. †Diacodon Cope 1875

†Diacodon alticuspis Cope 1875

†Diacodon puercensis Matthew 1918

G. †Ghambilentes Unknown 2000

†Ghambilentes foussanensis Gheerbrant and Hartenberger 1999

G. †Quadratodon De Bast and Smith 2017

†Quadratodon sigei De Bast and Smith 2017

G. †Seia Russell and Gingerich 1981

†Seia shahi Russell and Gingerich 1981

Fm. †Sespedectidae Novacek 1985

Subfm. †Sespedectinae Novacek 1985

G. †Colpocherus Beard and Dawson 2009

†Colpocherus mississippiensis Beard and Dawson 2009

G. †Crypholestes Novacek 1980

†Crypholestes vaughni Novacek 1976

Invalid names: Cryptolestes Novacek 1976 [replaced]

G. †Patriolestes Walsh 1998

†Patriolestes novaceki Walsh 1998

G. †Proterixoides Stock 1935

†Proterixoides davisi Stock 1935

G. †Sespedectes Stock 1935

†Sespedectes singularis Stock 1935

†Sespedectes stocki Novacek 1985

G. †Talpavoides Bown and Schankler 1982

†Talpavoides dartoni Bown and Schankler 1982

G. †Talpavus Marsh 1872

†Talpavus conjunctus Storer 1995

†Talpavus duplus Krishtalka 1976

†Talpavus sullivani Guthrie 1971

Fm. †Vastanidae Bajpai et al. 2005

G. †Vastania Bajpai et al. 2005

†Vastania sahnia Bajpai et al. 2005

Invalid names: Creotarsinae Hay 1930 [invalid subgroup]

G. †Ghamidtherium Sánchez-Villagra et al. 2007

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†Ghamidtherium dimaiensis Sánchez-Villagra et al. 2007

Subor. Soricomorpha Gregory 1910

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Fm. †Changlelestidae Tong and Wang 1993

G. †Changlelestes Tong and Wang 1993

†Changlelestes dissetiformis Tong and Wang 1993

G. †Ernosorex Wang and Li 1990

†Ernosorex jilinensis Wang and Li 1990

Subfm. †Tupaiodontinae Butler 1988

G. †Anatolechinos Wang 2008

†Anatolechinos huadianensis Wang and Li 1990

†Anatolechinos neimongolensis Wang 2008

G. †Ictopidium Zdansky 1930

†Ictopidium lechei Zdansky 1930

G. †Tupaiodon Matthew and Granger 1924

†Tupaiodon morrisi Matthew and Granger 1924

G. †Zaraalestes Storch and Dashzeveg 1997

†Zaraalestes minutus Matthew and Granger 1924

Invalid names: Ictopidium tatalgolensis Sulimski 1970 [synonym]

†Zaraalestes russelli Storch and Dashzeveg 1997

Fm. †Geolabididae McKenna 1960

G. †Batodonoides Novacek 1976

†Batodonoides powayensis Novacek 1976

†Batodonoides vanhouteni Bloch et al. 1998

†Batodonoides walshi Kelly 2010

G. †Centetodon Marsh 1872

†Centetodon altidens Marsh 1872

Invalid names: Didelphodus ventanus White 1952 [synonym]

†Centetodon aztecus Lillegraven et al. 1981

†Centetodon bacchanalis McGrew et al. 1959

†Centetodon bembicophagus Lillegraven et al. 1981

†Centetodon chadronensis Lillegraven et al. 1981

†Centetodon divaricatus Korth 1992

†Centetodon hendryi Lillegraven et al. 1981

†Centetodon kuenzii Lillegraven et al. 1981

†Centetodon magnus Clark 1936

†Centetodon marginalis Cope 1873

Invalid names: Embassis alternans Cope 1873 [synonym], Metacodon mellingeri Patterson and McGrew 1937 [synonym]

†Centetodon neashami Bown and Schankler 1982

†Centetodon patratus Bown and Schankler 1982

†Centetodon pulcher Marsh 1872

Invalid names: Hypacodon praecursor McKenna 1960 [synonym]

†Centetodon wolffi Macdonald 1965

Invalid names: Embassis Cope 1873 [synonym], Hypacodon McKenna 1960 [synonym], Protictops Peterson 1934 [synonym]

G. †Gobigeolabis Lopatin 2004

†Gobigeolabis verigranum Lopatin 2004

G. †Marsholestes McKenna 1992

†Marsholestes dasypelix Matthew 1909

G. †Stilpnodon Simpson 1935

†Stilpnodon simplicidens Simpson 1935

Invalid names: Centetodontinae Trouessart 1879 [synonym], Metacodontidae Butler 1948 [synonym], Protictops alticuspidens Peterson 1934 [nomen dubium]

G. †Nycticonodon Quinet 1964

Fm. †Plesiosoricidae Winge 1917

G. †Butselia Quinet and Misonne 1965

†Butselia biveri Quinet and Misonne 1965

G. †Ordolestes Lopatin 2006

†Ordolestes ordinatus Lopatin 2006

G. †Pakilestes Russell and Gingerich 1981

†Pakilestes lathrius Russell and Gingerich 1981

G. †Plesiosorex Pomel 1848

†Plesiosorex aydarlensis Kordikova 2000

†Plesiosorex coloradensis Wilson 1960

†Plesiosorex donroosai Green 1977

†Plesiosorex evolutus Ziegler 2006

†Plesiosorex germanicus Seemann 1938

†Plesiosorex greeni Martin and Lim 2004

†Plesiosorex latidens Hall 1929

†Plesiosorex schaffneri Engesser 1972

†Plesiosorex soricinoides Blainville 1838

Invalid names: Plesiosorex talpoides Pomel 1848 [synonym]

†Plesiosorex styriacus Hoffmann 1892

Invalid names: Meterix Hall 1929 [synonym]

G. †Pseudoneurogymnurus Gureev 1979

†Pseudoneurogymnurus shevyrevae Gureev 1979

†Pseudoneurogymnurus zhchikvadzei Gureev 1979

G. †Siamosorex Peigné et al. 2009

†Siamosorex debonisi Peigné et al. 2009

Invalid names: Butselidae Quinet and Misonne 1965 [synonym], Butseliinae Quinet and Misonne 1965 [empty], Plesiosoricinae Winge 1917 [empty]

Fm. Soricidae Fischer 1814 [shrew]

G. †Alloblarinella Storch 1995

†Alloblarinella europaea Reumer 1984

†Alloblarinella sinica Storch 1995

G. †Alluvisorex Hutchison 1966

†Alluvisorex arcadentes Hutchison 1966

†Alluvisorex chasseae Tedford 1961

G. †Asoriculus Kretzoi 1959

G. †Beremendia Kormos 1930

†Beremendia fissidens Petényi 1864

G. †Cokia Storch 1995

†Cokia kowalskae Storch 1995

†Cokia robusta Rzebik-Kowalska 1989

G. †Cretasorex Nessov and Gureyev 1981

†Cretasorex arkhangelskyi Nessov and Gureyev 1981

Subfm. Crocidosoricinae Reumer 1987

G. †Carposorex Crochet 1975

G. †Clapasorex Crochet 1975

†Clapasorex alvarezae van den Hoek Ostende 2003

Tr. †Crocidosoricini Reumer 1987

G. †Crocidosorex Lavocat 1951

G. †Miocrocidosorex Lopatin 2004

G. †Florinia Ziegler 1989

†Florinia stehlini Doben-Florin 1964

G. †Lartetium Ziegler 1989

†Lartetium petersbuchense Ziegler 1989

†Lartetium prevostianum Lartet 1851

G. †Miosorex Kretzoi 1959

†Miosorex grivensis Depéret 1892

†Miosorex pusilliformis Doben-Florin 1964

G. †Oligosorex Kretzoi 1959

†Oligosorex bruijni Gibert 1974

Tr. †Oligosoricini Gureev 1971

G. †Aralosorex Lopatin 2004

G. †Soricella Doben-Florin 1964

†Soricella discrepans Doben-Florin 1964

G. †Srinitium Hugueney 1976

†Srinitium caeruleum Ziegler 1998

†Srinitium marteli Hugueney 1976

G. †Taatsiinia Ziegler et al. 2007

†Taatsiinia hoeckorum Ziegler et al. 2007

G. †Tavoonyia Ziegler et al. 2007

†Tavoonyia altaica Ziegler et al. 2007

G. †Ulmensia Ziegler 1989

†Ulmensia antecedens Ziegler 1998

†Ulmensia ehrensteinensis Ziegler 1989

Subfm. Crocidurinae Milne Edwards 1874 [white-toothed shrew]

G. Crocidura Wagler 1832

Crocidura attila Dollman 1915

Crocidura giffardi de Winton 1898

Crocidura horsfieldi Tomes 1856 [Horsfield's shrew]

Crocidura kornfeldi Kormos 1934

Crocidura leucodon Hermann 1780

Crocidura martiensseni Neumann 1900

Crocidura olivieri Lesson 1827 [African giant shrew]

Crocidura poensis Fraser 1843

Crocidura russula Herrmann 1780 [Greater white-toothed shrew]

Crocidura suaveolens Pallas 1811

Invalid names: Crocidura mimula Miller 1901 [synonym]

Crocidura sururae Heller 1910

Crocidura zimmermanni Wettstein 1953

Sorex (Crocidura) caeruleus Kerr 1792

Invalid names: Sorex caerulescens Shaw 1800 [synonym]

Invalid names: Heliosorex Heller 1910 [synonym], Leucodon Fatio 1869 [synonym], Paurodus Schulze 1897 [synonym], Rhinomus Murray 1861 [synonym]

Tr. Crocidurini Milne Edwards 1872

G. Diplomesodon Brandt 1952

G. †Feroculus Kelaart 1852

G. Myosorex Gray 1838

Invalid names: Congosorex Heim de Balsac and Lamotte 1956 [synonym]

G. †Paracrocidura Heim de Balsac 1956

G. †Praesorex Thomas 1913

G. †Solisorex Thomas 1924

G. †Surdisorex Thomas 1906

G. Sylvisorex Thomas 1904

Tr. †Scutisoricini Allen 1917

G. †Scutisorex Thomas 1913

G. Suncus Ehrenberg 1832

Suncus etruscus Savi 1822

Suncus madagascariensis Coquerel 1848 [Madagascan pygmy shrew]

Suncus murinus Linnaeus 1766 [Asian house shrew]

Suncus varilla Thomas 1895 [Lesser dwarf shrew]

Invalid names: Suncus orangiae Roberts 1924 [synonym]

Invalid names: Pachyura Selys Longchamps 1839 [synonym], Plerodus Schulze 1897 [synonym]

Subfm. †Heterosoricinae Viret and Zapfe 1951

G. †Atasorex Lopatin 2004

†Atasorex edax Lopatin 2004

G. †Dinosorex Engesser 1972

†Dinosorex anatolicus van den Hoek Ostende 1995

†Dinosorex engesseri Rabeder 1998

†Dinosorex huerzeleri Engesser 1975

†Dinosorex pachygnathus Engesser 1972

†Dinosorex sansaniensis Lartet 1851

†Dinosorex zapfei Engesser 1975

G. †Domnina Cope 1873

†Domnina crassigenis Cope 1873

†Domnina dakotensis Macdonald 1970

†Domnina gradata Cope 1873

†Domnina greeni Macdonald 1963

†Domnina sagittariensis Kihm and Schumaker 2008

†Domnina thompsoni Simpson 1941

Invalid names: Miothen Cope 1873 [synonym]

G. †Gobisorex Sulimski 1970

†Gobisorex akhmetievi Lopatin 2004

†Gobisorex kingae Sulimski 1970

G. †Heterosorex Gaillard 1915

†Heterosorex ruemkeae Doukas 1986

†Heterosorex wangi Storch and Qiu 1991

G. †Ingentisorex Hutchison 1966

†Ingentisorex tumididens Hutchison 1966

G. †Lusorex Storch and Qiu 2004

†Lusorex taishanensis Storch and Qiu 2004

G. †Noritrimylus Korth et al. 2022

†Noritrimylus compressus Galbreath 1953

†Noritrimylus dakotensis Repenning 1967

†Noritrimylus metaxy Korth 2020

G. †Paradomnina Hutchison 1966

†Paradomnina relictus Hutchison 1966

G. †Pseudotrimylus Gureev 1971

†Pseudotrimylus blacki Martin and Lim 2004

†Pseudotrimylus mawbyi Repenning 1967

†Pseudotrimylus roperi Wilson 1960

Invalid names: Trimylus Roger 1885 [synonym]

G. †Quercysorex Engesser 1975

†Quercysorex herrlingensis Palmowski and Wachendorf 1966

†Quercysorex primaevus Filhol 1884

†Quercysorex ulmensis Ziegler 1998

G. †Wilsonosorex Martin 1978

†Wilsonosorex bateslandensis Martin 1978

†Wilsonosorex conulatus Martin 1978

Invalid names: Sorex kinkelini Zinndorf 1901 [nomen dubium]

Subfm. †Limnoecinae Repenning 1967

G. †Angustidens Repenning 1967

†Angustidens excultus Mayr and Fahlbusch 1975

†Angustidens vireti Wilson 1960

G. †Limnoecus Stirton 1930

†Limnoecus compressus Wilson 1960

†Limnoecus niobrarensis Macdonald 1947

†Limnoecus tricuspis Stirton 1930

†Limnoecus truyolsi Gibert 1974

G. †Lunanosorex Jin and Kawamura 1996

Lunanosorex lii Jin and Kawamura 1996

G. †Nesiotites Bate 1945

†Nesiotites corsicanus Bate 1945

†Nesiotites hidalgo Bate 1945

†Nesiotites rafelinensis Rofes et al. 2012

†Nesiotites similis Hensel 1855

G. †Paenepetenyia Storch 1995

†Paenepetenyia zhudingi Storch 1995

G. †Parydrosorex Wilson 1968

†Parydrosorex concisus Wilson 1968

G. †Petenyiella Kretzoi 1956

†Petenyiella zelcea Sulimski 1959

Invalid names: Allopachyura Kormos 1934 [synonym]

G. †Podihik Deraniyagala 1958

G. †Protosorex Scott 1894

†Protosorex crassus Scott 1894

G. †Siwalikosorex Sahni and Khare 1976

Subfm. Soricinae Fischer von Waldheim 1817 [red-toothed shrew]

Tr. †Allosoricini Fejfar 1966

G. †Allosorex Fejfar 1966

G. †Paenelimnoecus Baudelot 1972

Tr. †Amblycoptini Kormos 1926

G. †Kordosia Mészáros 1997

Tr. Anourosoricini Anderson 1879

G. †Paranourosorex Rzebik-Kowalska 1975

Tr. Blarinini Kretzoi 1965

G. †Adeloblarina Repenning 1967

G. Blarina Gray 1838 [American short-tailed shrew]

Invalid names: Anotus Wagner 1855 [synonym], Mamblarinaus Herrera 1899 [synonym], Talposorex Pomel 1848 [synonym]

G. †Blarinoides Sulimski 1959

G. Cryptotis Pomel 1848 [small-eared shrew]

Invalid names: Brachysorex Duvernoy 1842 [synonym], Soriciscus Coues 1877 [synonym]

G. †Mafia Reumer 1984

G. †Paracryptotis Hibbard 1950

G. †Peisorex Kowalski and Li 1963

G. †Shikamainosorex Hasegawa 1957

G. †Sulimskia Reumer 1984

G. †Tregosorex Hibbard and Jammot 1971

G. †Hemisorex Baudelot 1967

Tr. †Nectogalini Anderson 1879

G. †Arctisorex Hutchison and Harington 2002

G. †Beckiasorex Dalquest 1972

G. †Crusafontina Gibert 1975

Invalid names: Anouroneomys Brown 1980 [synonym]

G. †Hesperosorex Hibbard 1957

G. Notiosorex Baird 1877 [gray shrew]

Tr. Neomyini Matschie 1909

G. †Amblycoptus Kormos 1926

G. Anourosorex Milne-Edwards 1872 [mole shrew]

Invalid names: Pygmura Anderson 1875 [synonym]

G. †Builstynia Ziegler et al. 2007

G. Chimarrogale Anderson 1877 [Asiatic water shrew]

Invalid names: Crossogale Thomas 1921 [synonym]

G. Chodsigoa Kastchenko 1907

G. Episoriculus Ellerman and Morrison-Scott 1951

G. Megasorex Hibbard 1950

G. Nectogale Milne-Edwards 1870

G. Neomys Kaup 1829 [water shrew]

Invalid names: Crossopus Wagler 1832 [synonym], Hydrogale Kaump 1829 [synonym], Leucorrhynchus Kaup 1829 [synonym], Myosictis Pomel 1854 [synonym], Pinalia Gray 1838 [synonym]

G. †Neomysorex Rzebik-Kowalska 1981

G. Soriculus Blyth 1845

Invalid names: Amblycoptinae Kormos 1926 [synonym], Anourosoricinae Anderson 1879 [synonym], Crossopinae Milne Edwards 1872 [synonym], Hydrosoridae Anon 1838 [synonym], Nectogalinae Anderson 1879 [synonym], Soriculi Winge 1917 [synonym]

Tr. Soricini Fischer von Waldheim 1817

G. †Anchiblarinella Hibbard and Jammot 1971

G. †Antesorex Repenning 1967

G. Blarinella Thomas 1911 [Asiatic short-tailed shrew]

G. †Deinsdorfia Heller 1963

G. †Drepanosorex Kretzoi 1941

G. †Paenesorex Ziegler 2003

G. †Petenyia Kormos 1934

G. †Planisorex Skinner and Hibbard 1972

G. Sorex Linnaeus 1758 [long-tailed shrew]

Invalid names: Amphisorex Duvernoy 1835 [synonym], Atophyrax Merriam 1884 [synonym], Corsira [synonym], Homalurus Schulze 1890 [synonym], Hydrogale Pomel 1848 [synonym], Microsorex Baird 1877 [synonym], Musaraneus Brisson 1762 [synonym], Neosorex Baird 1857 [synonym], Otisorex De Kay 1842 [synonym], Soricidus Altobello 1927 [synonym]

Subfm. †Soricolestinae Lopatin 2002

G. †Soricolestes Lopatin 2002

†Soricolestes soricavus Lopatin 2002

G. †Zelceina Sulimski 1962

†Zelceina kormosi Schlosser 1924

†Zelceina podlesicensis Rzebik-Kowalska 1990

†Zelceina soriculoides Sulimski 1959

Invalid names: Allosoricinae Fejfar 1966 [empty]

Superfm. Soricoidea Gill 1872

Fm. †Apternodontidae Matthew 1910

G. †Apternodus Matthew 1903

†Apternodus baladontus Asher 2002

†Apternodus brevirostris Schlaikjer 1934

†Apternodus dasophylakas Asher 2002

†Apternodus gregoryi Schlaikjer 1933

†Apternodus iliffensis Galbreath 1953

†Apternodus major Asher 2002

†Apternodus mediaevus Matthew 1903

Subfm. †Asiapternodontinae Lopatin 2006

G. †Asiapternodus Lopatin 2003

G. †Iconapternodus Tong 1997

†Iconapternodus qii Tong 1997

Invalid names: Apternodontinae Matthew 1910 [empty], Oligoryctinae Asher et al. 2002 [empty], Parapternodontinae Asher et al. 2002 [empty]

Fm. †Micropternodontidae

G. †Clinopternodus Clark 1937

†Clinopternodus gracilis Clark 1936

Invalid names: Clinodon Clark 1936 [synonym]

G. †Cryptoryctes Reed 1954

†Cryptoryctes kayi Reed 1954

G. †Micropternodus Matthew 1903

†Micropternodus borealis Matthew 1903

†Micropternodus montrosensis Ostrander 1983

†Micropternodus morgani Stirton and Rensberger 1964

†Micropternodus strophensis White 1954

Invalid names: Kentrogomphios White 1954 [synonym]

G. †Sinosinopa Qi 1987

†Sinosinopa sinensis Qi 1987

Fm. †Nyctitheriidae Simpson 1928

Subfm. †Amphidozotheriinae Sige 1976

G. †Darbonetus Crochet 1974

Subfm. †Asionyctiinae Missiaen and Smith 2005

G. †Asionyctia Missiaen and Smith 2005

G. †Bayanulanius Meng et al. 1998

G. †Edzenius Lopatin 2006

G. †Oedolius Russell and Dashzeveg 1986

Subfm. †Eosoricodontinae Lopatin 2005

G. †Eosoricodon Lopatin 2005

G. †Indonyctia Das et al. 2022

†Indonyctia cambayensis Das et al. 2022

G. †Leptacodon Matthew and Granger 1921

†Leptacodon acherontus Secord 2008

†Leptacodon catulus Krishtalka 1976

†Leptacodon choristus Secord 2008

†Leptacodon donkroni Rose et al. 2012

†Leptacodon minutulus Dorr 1958

†Leptacodon munusculum Simpson 1935

†Leptacodon nascimentoi Estravís 1996

†Leptacodon packi Jepsen 1930

†Leptacodon proserpinae Van Valen 1978

†Leptacodon tener Matthew and Granger 1921

G. †Limaconyssus Gingerich 1987

†Limaconyssus habrus Gingerich 1987

G. †Myolestes Matthew 1909

Subfm. †Nyctitheriinae Simpson 1928

G. †Cryptotopos Crochet 1974

G. †Euronyctia Sigé 1997

G. †Oligonyctia Smith 2004

G. †Plagioctenodon Bown 1979

G. †Saturninia Stehlin 1940

G. †Scraeva Cray 1973

Invalid names: Arvaldus Cray 1973 [synonym]

G. †Sigenyctia Smith 2006

G. †Nyctitherium Marsh 1872

†Nyctitherium celatus Cope 1875

Invalid names: Anaptomorphus minimus Loomis 1906 [synonym]

†Nyctitherium christopheri Krishtalka and Setoguchi 1977

†Nyctitherium curtidens Matthew 1909

†Nyctitherium gunnelli Murphey and Kelly 2017

†Nyctitherium nitidum Marsh 1872

†Nyctitherium priscum Marsh 1872

†Nyctitherium priscus Marsh 1872

†Nyctitherium serotinum Marsh 1872

†Nyctitherium velox Marsh 1872

Invalid names: Nyctilestes Marsh 1872 [synonym]

Subfm. †Placentidentinae Russell et al. 1973

G. †Ceutholestes Rose and Gingerich 1987

G. †Placentidens Russell et al. 1973

G. †Plagioctenoides Bown 1979

G. †Pontifactor West 1974

G. †Remiculus Russell 1964

G. †Wyonycteris Gingerich 1987

Subfm. †Praolestinae Lopatin 2006

G. †Bumbanius Russell and Dashzeveg 1986

G. †Praolestes Matthew et al. 1929

G. †Protentomodon Simpson 1928

†Protentomodon ursirivalis Simpson 1928

Subfm. †Talpilestinae Tong and Wang 2006

G. †Talpilestes Tong and Wang 2006

G. †Yuanqulestes Tong 1997

†Yuanqulestes qiui Tong 1997

Invalid names: Paradoxonycteris soricodon Revilliod 1919 [nomen dubium]

Fm. †Oligoryctidae Asher et al. 2002

G. †Oligoryctes Hough 1956

†Oligoryctes altitalonidus Clark 1936

†Oligoryctes cameronensis Hough 1956

†Oligoryctes tenutalonidus Korth et al. 2022

Fm. †Parapternodontidae Asher et al. 2002

G. †Koniaryctes Robinson and Kron 1998

†Koniaryctes paulus Robinson and Kron 1998

G. †Parapternodus Bown and Schankler 1982

†Parapternodus antiquus Bown and Schankler 1982

Infraor. †Soricota Kalandadze and Rautian 1992

Superfm. †Nesophontoidea Anthony 1916

Fm. †Nesophontidae

G. †Nesophontes Anthony 1916

Superfm. Talpoidea Fischer de Waldheim 1817

Fm. †Proscalopidae Reed 1961

G. †Mesoscalops Reed 1960

G. †Oligoscalops Reed 1961

G. †Proscalops Matthew 1901

Fm. Talpidae Fischer de Waldheim 1817 [mole]

G. †Anomodon Leconte 1848

G. †Asthenoscapter Hutchison 1974

G. Desmana Guldenstaedt 1777 [Russian desman]

Invalid names: Caprios Wagler 1830 [synonym], Desmagale Kretzoi 1953 [synonym], Desman Lacepede 1799 [synonym], Desmanus Rafinesque 1815 [synonym], Mygale Cuvier 1800 [synonym], Myogale Brandt 1836 [synonym], Myogalea Fischer 1829 [synonym], Palaeospalax Owen 1846 [synonym]

Subfm. Desmaninae Thomas 1912

Invalid names: Myogalina Mivart 1871 [synonym]

G. †Desmanodon Engesser 1980

G. †Domninoides Green 1956

G. †Entomodon Marsh 1872

G. †Eotalpa Sigé et al. 1977

G. †Galeospalax Pomel 1848

G. †Geolabis Cope 1884

Invalid names: Metacodon Clark 1936 [synonym]

G. †Hugueneya van den Hoek Ostende 1989

G. †Hyporyssus Pomel 1848

G. †Mioscalops Ostrander et al. 1986

G. Mogera Pomel 1848

G. †Mongolopala Ziegler et al. 2007

G. Neurotrichus Günther 1880 [shrew mole]

G. †Oreotalpa Lloyd and Eberle 2008

G. Parascalops True 1894

G. Parascaptor Gill 1875

Subfm. †Proscalopinae Reed 1961

Invalid names: Arctoryctinae Reed and Turnbull 1965 [synonym]

G. †Quadrodens Macdonald 1970

Invalid names: Palaeoscalopus Macdonald 1970 [synonym]

G. Scaptochirus Milne-Edwards 1867 [Short-faced mole]

Invalid names: Chiroscaptor Heude 1898 [synonym]

G. †Scaptogale Trouessart 1898

Invalid names: Echinogale Pomel 1848 [synonym]

Subfm. †Suleimaninae Van den Hoek Ostende 2001

G. Talpa Linnaeus 1758

Invalid names: Asioscalops Stroganov 1941 [synonym], Eoscalops Stroganov 1941 [synonym], Euroscaptor Miller 1940 [synonym], Heterotalpa Peters 1863 [synonym], Talpops Gervais 1868 [synonym]

Subfm. Talpinae Fisher von Waldheim 1817

G. †Tegulariscaptor Sansalone et al. 2017

Subfm. Uropsilinae Dobson 1883

Invalid names: Chrysochlorina Gray 1825 [empty], Myaladae Gray 1821 [synonym], Scalopinae Gill 1875 [empty], Soricina Gray 1825 [empty], Talpina Gray 1825 [empty], Tenrecina Gray 1825 [empty], Tupaina Gray 1825 [empty]

Invalid names: Micropternodontoidea Stirton and Rensberger 1964 [empty], Solenodontoidea Gill 1972 [empty]

Invalid names: Lipotyphla Haeckel 1866 [synonym]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

phosphaticsubp

Maximum body mass:

10 gs

Minimum body mass:

10 gs

Body mass estimate:

10 g

Environment:

terrestrialsubc

Locomotion:

actively mobilec

Life habit:

scansorialsubc

Diet:

insectivoresubc

Reproduction:

viviparoussubc

Created:

2005-09-06 20:43:42

Modified:

2005-09-06 22:44:32

Source: s = species, subc = subclass, c = class, subp = subphylum

References: Legendre 1988, Carroll 1988, Ji et al. 2002, Lillegraven 1979, Hendy et al. 2009

Age range

Maximum range based only on fossils: base of the Middle Turonian to the top of the Holocene or 93.50000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 89.3 Ma

Collections (2191 total)

Time interval	Ma	Country or state	Original ID and collection number
Oldest occurrences
Middle Turonian - Late/Upper Turonian	93.5 - 89.3	Uzbekistan (Navoi)	Cretasorex arkhangelskyi (58179)
Judithian	83.5 - 70.6	Canada (Alberta)	Nyctitheriidae indet. (14451)
Maastrichtian	72.1 - 66.0	Portugal (Coimbra)	Leptacodon sp. (49276)
Puercan	66.0 - 63.3	USA (New Mexico)	Erinaceomorpha indet. (14760 14794)
Puercan	66.0 - 63.3	USA (Montana)	Leptacodon proserpinae (14766)
Puercan	66.0 - 63.3	USA (Utah)	Propalaeosinopa sp. (14790)
Danian	66.0 - 61.6	Belgium (Hainaut)	Quadratodon sigei (132949)
Danian	66.0 - 61.6	USA (North Dakota)	Leptacodon tener, Propalaeosinopa albertensis (14962)
To2	63.8 - 60.9	Canada (Alberta)	Leptacodon munusculum, Leptacodon tener, Nyctitheriinae indet., Propalaeosinopa albertensis, Bessoecetor sp. 1, Bessoecetor sp. 2, Propalaeosinopa septentrionalis (14894)
Torrejonian	63.8 - 60.9	USA (Montana)	Bessoecetor diluculi (14913) Pantomimus leari (14918) Stilpnodon simplicidens, Leptacodon munusculum, Leptonysson basiliscus, Propalaeosinopa diluculi (14906)
Torrejonian	63.8 - 60.9	USA (New Mexico)	Pentacodon inversus (14832 14865 14873) Pentacodon inversus, Prodiacodon puercensis (14815) Pentacodon occultus, Prodiacodon puercensis (14833 14893) Pentacodon sp. (14819) Pentacodon sp., Prodiacodon sp. (14816) Prodiacodon puercensis (14859)
Torrejonian	63.8 - 60.9	Canada (Alberta)	Bessoecetor septentrionalis (143199 143200) Propalaeosinopa sp., Lipotyphla indet. (14828)
Torrejonian	63.8 - 60.9	USA (Utah)	Pantolestidae indet. (14830)
Torrejonian	63.8 - 60.9	USA (Wyoming)	Leptacodon munusculum (14912) Leptacodon tener, Pantolestinae indet., Prodiacodon puercensis, Propalaeosinopa diluculi (14886) Propalaeosinopa diluculi (14898 14900 14904)
Ti1	61.7 - 56.8	Canada (Alberta)	Leptacodon munusculum, Leptacodon packi, Leptacodon tener, Limaconyssus sp., Nyctitherium sp., Nyctitheriidae indet., Prodiacodon puercensis, Propalaeosinopa septentrionalis (14939) Propalaeosinopa septentrionalis (14938)
Ti3	61.7 - 56.8	Canada (Alberta)	Leptacodon munusculum, Leptacodon tener, Bessoecetor septentrionalis (14928) Leptacodon packi, Litolestes ignotus, Erinaceidae indet., Propalaeosinopa albertensis, Propalaeosinopa septentrionalis (14992)
Ti4	61.7 - 56.8	Canada (Saskatchewan)	Nyctitherium sp., Oncocherus krishtalkai, Litolestes ignotus, Propalaeosinopa sp., Bessoecetor pilodontus, Bessoecetor krausei, Palaeosinopa reclusum, Aatotomus placochton, Lipotyphla indet., Leptacodon packi (14978)
Tiffanian	61.7 - 56.8	Canada (Alberta)	Leptacodon packi, Erinaceidae indet. A, Erinaceidae indet. B (14929) Leptacodon packi, Leptacodon tener, Nyctitherium sp., Litolestes sp., Propalaeosinopa diluculi (14973) Leptacodon packi, Litolestes ignotus (14934) Leptacodon packi, Propalaeosinopa septentrionalis (14954) Leptacodon tener, Nyctitheriidae indet., Oncocherus krishtalkai, Litolestes ignotus, Propalaeosinopa sp., Pantolestidae indet., Eudaemonema cuspidata (14995) Nyctitheriidae indet., Erinaceidae indet. (14951) Oncocherus krishtalkai, Litolestes ignotus (14941) Propalaeosinopa albertensis (14949 14957) Propalaeosinopa albertensis, Propalaeosinopa septentrionalis (14933) Stilpnodon sp., Litolestes ignotus, Erinaceidae indet., Propalaeosinopa albertensis, Lipotyphla indet., Propalaeosinopa septentrionalis (14959)
Tiffanian	61.7 - 56.8	USA (Wyoming)	Apternodontidae indet., Leptacodon packi, Wyonycteris galensis, Litolestes ignotus (15049) Bessoecetor sp., Prodiacodon puercensis (198262) Cedrocherus aceratus, Leptacodon munusculus, Propalaeosinopa diluculi (14936) Leptacodon acherontus, Leptacodon munusculum, Leptacodon tener, Cedrocherus ryani, Leptonysson orthius, Propalaeosinopa diluculi (15017) Leptacodon packi, Litolestes ignotus, Pantolestidae indet. (15032) Leptacodon packi, Plagioctenodon sp., Leptacodon sp., Litolestes sp. (26569) Leptacodon sp. (14926) Leptacodon sp., Propalaeosinopa thomsoni (15034) Leptacodon tener (14974) Leptacodon tener, Palaeosinopa sp., Talpavoides dartoni, Propalaeosinopa diluculi (14952) Litolestes ignotus (15057) Palaeosinopa simpsoni (14944) Pentacodon sp. (15067) Prodiacodon puercensis (198264) Propalaeosinopa diluculi (14987 15014) Wyonycteris microtis, Litolestes ignotus (15065)
Tiffanian	61.7 - 56.8	USA (North Dakota)	Leptacodon sp., Entomolestes sp., Propalaeosinopa sp. (15000) Leptacodon tener, Propalaeosinopa albertensis (14931) Pantolestoidea indet. (14946) Propalaeosinopa albertensis (14977) Propalaeosinopa diluculi (15006)