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Zolhafah bella

Reptilia - Testudines

Taxonomy
Zolhafah bella was named by Lapparent de Broin and Werner (1998). Its type specimen is TUB Vb-173, a partial skull (partial skull without mandibles), and it is a 3D body fossil. Its type locality is Ammonite Hill interdunal channel 31 (271080/1), which is in a Maastrichtian coastal sandstone/claystone in the Dakhla Formation of Egypt.

Synonymy list
YearName and author
1998Zolhafah bella Lapparent de Broin and Werner p. 138 figs. 3a-b; plates 1-2
2006Zolhafah bella Gaffney et al. p. 70
2012Zolhafah bella Rabi et al. p. 672

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
RankNameAuthor
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
Sauropsida
classReptilia
Testudinata(Oppel 1811)
orderTestudinesBatsch 1788
suborderPleurodira
Bothremydidae(Baur 1891)
tribeBothremydini
subtribeBothremydina
genusZolhafah
speciesbella

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Zolhafah bella Lapparent de Broin and Werner 1998
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Diagnosis
ReferenceDiagnosis
F. Lapparent de Broin and C. Werner 1998Bothremydid belonging to the Bothretnys group by the presence of pits in the posterior part of the enlarged triturating skull surfaces, in the maxilla, the jugal and the palatine. Zolhafah bella n.g., n.sp., differs from Bothremys cooki Leidy, 1865 and Rosasia Carrington Da Costa, 1940, principally by its more rounded snout, absolutely not medio-anteriorly prominent, its more shortened posterior skull part, its basioccipital and basisphenoid wider and its basiphenoid shorter than in both others, from Rosasia, at least, by its flat posterior palate (absence of podocnemidoid fossa) and: as in B. cooki, the choanae are at the level of the triturating pits (apomorphic characters). Its triturating pits are intermediate in depth between Rosasia and B. cooki. Its external nostril is much larger, higher and wider (plesiomorphic) and it is more vertical than in B. cooki. Moreover, Zolhafah bella n.g., n.sp., differs from Bothremys cooki and Rosasia by many details such as its frontal being longer than the prefrontal, its elongated orbits smaller than in Rosasia, larger than in B. cooki (where they are subcircular) and its higher anterior premaxillary surface (plesiomorphic characters), its maxilla more developed below the orbit than in B. cooki (in the latter. the orbit is higher than in Rosasia) and its ventral premaxillary-maxillary border thickened as in Rosasia, but still thicker than in Rosasia (apomorphic) (thin in B. cooki). Its anterior palatine sulcus between the triturating surfaces is shorter (apomorphic) and wider (plesiomorphic) than in both others. Its palatino-pterygoid suture is relatively wide, as in B. cooki, compared to Rosasia where it is narrowed. The occipital condyle was slightly posterior to the articular processeso f the quadrates,p robably as in Rosasiaa nd B. woki, and located less forward than in B. harheri. Size of Rosasia and B. cooki, much smaller than B. hurheri (Schmidt. 1940).
E. S. Gaffney et al. 2006[as for genus Zolhafah]
Measurements
No measurements are available
Composition: phosphaticsubp
Environment: freshwatersubo
Locomotion: actively mobilec
Life habit: amphibioussubo
Diet: omnivoresubo
Reproduction: oviparoussubo
Created: 2005-08-26 14:50:50
Modified: 2005-08-26 16:50:50
Source: subo = suborder, c = class, subp = subphylum
References: Uetz 2005, Hendy et al. 2009, Carroll 1988

Age range: Maastrichtian or 72.20000 to 66.00000 Ma

Collections: one only


Time interval Ma Country or state Original ID and collection number
Maastrichtian72.2 - 66.0Egypt Zolhafah bella (type locality: 83310)