Bore Kevelaer, depth 70m (Miocene of Germany)

Where: North Rhein-Westfalia, Germany (51.6° N, 6.3° E)

• Paleocoordinates: 49.0° N, 8.5° E (Wright 2013)

• coordinate estimated from map

• small collection-level geographic resolution

When: Middle Miocene (16.0 - 11.6 Ma)

• Reinbek-Bislicher stage or Reinbek-Tichelh. stage

• bed-level stratigraphic resolution

Environment/lithology: marine; unlithified, fine-grained, brown sandstone

Size classes: macrofossils, mesofossils, microfossils

Preservation: original aragonite

Collection methods: core, mechanical, peroxide, sieve,

• well drilling, 1994

Primary reference: G. Wienrich. 1997. Die Fauna des marinen Miozäns von Kevelaer (Niederrhein). Band 1 Foraminiferen, Anthozoen. Backhuys Publishers, Leiden 1:1-187 [M. Aberhan/S. Nurnberg/S. Nurnberg]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 62887: authorized by Martin Aberhan, entered by Sabine Nürnberg on 01.08.2006

Creative Commons license: CC0 (CC0)

Taxonomic list

Show authors, comments, and common names
Globothalamea
 Rotaliida - Nonionidae
Nonion boueanum, Nonion affine, Nonion perfossum, Cribrononion subnodosum
 Rotaliida - Cibicididae
Planulina austriaca, Cibicides dutemplei, Cibicides lobatulus, Cibicides pseudoungerianus
Cibicides dutemplei dOrbigny 1846
Cibicides lobatulus Walker and Jacob 1798
incl. 2 specimens C. lobatulus var. ornatus
Foraminifera
 Lituolacea - Ataxophragmiidae
Martinottiella communis
Martinottiella communis Huang et al. 2019
 Buliminoidea - Buliminidae
Virgulina schreibersiana, Virgulina (Virgulinella) pertusa
 Asterigerinacea - Asterigerinidae
Asterigerina guerichi
Asterigerina guerichi
A. gürichi staeschi
 Discorbacea - Bagginidae
Cancris auriculus
Cancris auriculus Fichtel and Moll 1798
 Buliminacea - Uvigerinidae
Uvigerina semiornata
Uvigerina semiornata
U. semiornata for. semiornata
 Rotalioidea - Elphidiidae
Elphidium inflatum, Elphidium hauerinum
 Lituolida - Lituolidae
Bigenerina nodosaria, Textularia gramen
Textularia gramen dOrbigny 1846
Tubothalamea
 Miliolida - Miliolidae
Sigmoilina tenuis
Nodosariata
 Nodosariida - Nodosariidae
Nodosaria bacillum, Nodosaria longiscata, Lagena cf. emaciata, Lagena sulcata, Lagena globosa, Dentalina intermittens, Dentalina bifurcata, Dentalina grossicosta, Dentalina acuticauda, Dentalina konincki, Dentalina pauperata
Nodosaria longiscata d'Orbigny 1846
Lagena sulcata Williamson 1848
Lagena globosa Montagu 1803
Dentalina konincki Reuss 1861
Dentalina pauperata d'Orbigny 1846
 Polymorphinida - Polymorphinidae
Guttulina problema, Globulina gibba, Polymorphina charlottensis, Pseudopolymorphina subnodosa, Sigmomorphina schwageri
Guttulina problema d'Orbigny 1826
Globulina gibba d'Orbigny 1832
incl. 1 specimen G. gibba for. punctata and 1 specimen G. gibba rugosa
 Vaginulinida - Vaginulinidae
Lenticulina (Lenticulina) calcar, Lenticulina (Lenticulina) inornata, Lenticulina (Robulus) cultrata, Lenticulina (Robulus) similis, Lenticulina (Robulus) ornata
 Robertinida - Ceratobuliminidae
Ceratobulimina haueri
Foraminifera
 Spiroplectamminacea - Spiroplectamminidae
Spiroplectammina carinata
Anthozoa
 Scleractinia - Caryophylliidae
Deltocyathus reussi
 Scleractinia - Flabellidae
Flabellum sp., Flabellum tuberculatum, Flabellum cristatum, Flabellum pompeckji
Flabellum sp. Lesson 1831 stony coral
Flabellum tuberculatum Keferstein 1859 stony coral
Flabellum cristatum Milne-Edwards and Haime 1851 stony coral
Flabellum pompeckji Krejci 1926 stony coral
 Scleractinia - Micrabaciidae
Stephanophyllia imperialis
Stephanophyllia imperialis Michelin 1841 stony coral
Gymnolaemata
 Cheilostomata - Cupuladriidae
Reussirella haidingeri1
 Cheilostomata - Lunulitidae
Lunulites voigti1, Lunulites burdigalensis1
Sedentaria
 Sabellida - Serpulidae
Serpulidae indet.1
Serpulidae indet.1 Rafinesque 1835
Serpulidae indet. D
Bivalvia
 Nuculanida - Yoldiidae
Yoldia (Yoldia) glaberrima1, "Portlandia (Yoldiella) pygmaea" = Yoldiella1
Yoldia (Yoldia) glaberrima1 von Münster 1837 clam
"Portlandia (Yoldiella) pygmaea" = Yoldiella1 clam
 Nuculanida - Nuculanidae
"Nuculana (Saccella) westendorpi" = Saccella westendorpi1
"Nuculana (Saccella) westendorpi" = Saccella westendorpi1 Nyst and Westendorp 1839 pointed nut clam
 Nuculida - Nuculidae
Nuculoma haesendoncki1
Nuculoma haesendoncki1 nut clam
N. haesendoncki hanseata
 Arcida - Limopsidae
"Limopsis (Pectunculina) anomala" = Limopsis1, Limopsis (Pectunculina) lamellata1, Limopsis (Limopsis) aurita1
"Limopsis (Pectunculina) anomala" = Limopsis1 clam
Limopsis (Limopsis) aurita1 Brocchi 1814 clam
 Arcida - Arcidae
Scapharca (Scapharca) diluvii1
 Pectinida - Pectinoidae
Korobkovia woodi1, Pseudamussium lilli1, Aequipecten seniensis1
Pseudamussium lilli1 Pusch 1837 scallop
 Ostreida - Pinnidae
Atrina pectinata1
Atrina pectinata1 Linnaeus 1758 oyster
 Carditida - Carditidae
Cyclocardia sp.1
Cyclocardia sp.1 Conrad 1867 clam
 Carditida - Astartidae
Astarte (Astarte) radiata1, Goodallia (Goodallia) angulata1
Astarte (Astarte) radiata1 Nyst and Westendorp 1839 clam
Astarte radiata fa. radiata 28 specimens, Astarte radiata fa. convexior 33 specimens, Astarte radiata fa. goldfussi 2 specimens
Goodallia (Goodallia) angulata1 Lehmann 1885 clam
 Lucinida - Lucinidae
"Cavilucina (Goniomyrtea) droueti" = Gonimyrtea droueti1
"Cavilucina (Goniomyrtea) droueti" = Gonimyrtea droueti1 Nyst 1861 clam
C. droueti droueti
 Pandorida - Lyonsiidae
Lyonsia (Lyonsia) aff. norwegica1
 Pholadida - Corbulidae
Corbula (Varicorbula) gibba1
Corbula (Varicorbula) gibba1 Oliva 1792 clam
C. (V.) gibba gibba
 Cardiida - Cardiidae
Parvicardium straeleni1, Parvicardium scabrum1, Habecardium dingdense1
Parvicardium scabrum1 Philippi 1844 cockle
Habecardium dingdense1 Lehmann 1885 cockle
 Cardiida - Tellinidae
"Macoma elliptica" = Laciolina magna1
"Macoma elliptica" = Laciolina magna1 Spengler 1798 tellin clam
 Cardiida - Semelidae
sensu lato Abra lehmanni1
 Cardiida - Glossidae
Glossus (Glossus) lunulatus1
 Cardiida - Mactridae
Spisula elliptica1
Spisula elliptica1 Brown 1827 clam
 Cardiida - Veneridae
"Gouldia minima" = Gouldia (Gouldia) minima1, Venus (Ventricoloidea) multilamella1
"Gouldia minima" = Gouldia (Gouldia) minima1 Montagu 1803 venus clam
Venus (Ventricoloidea) multilamella1 Lamarck 1818 venus clam
V. (V.) multilamella multilamella
Scaphopoda
 Dentaliida - Dentaliidae
Lentigodentalium pseudomutabile1, "Pseudantalis aturensis" = Dentalium (Pseudantilis)1
"Pseudantalis aturensis" = Dentalium (Pseudantilis)1 tusk shell
 Gadilida - Gadilidae
Gadila dingdenensis1
Gastropoda
 Heterostropha - Acteonidae
Acteon (Acteon) semistriatus3, Crenilabium terebelloides3
Acteon (Acteon) semistriatus3 snail
spelled as 'Actaeon'
 Heterostropha - Ringiculidae
Ringicula (Ringiculina) buccinea3
 Heterostropha - Pyramidellidae
"Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros3, "Turbonilla (Pyrgolampros) undulata" = Pyrgolampros3, Pyramidella (Voluspa) plicosa3, Syrnola hoernesi3, Turbonilla (Turbonilla) gastaldi3, Turbonilla (Turbonilla) koeneniana3, Eulimella (Eulimella) acicula3, Odostomia (Brachystomia) sp.3, Odostomia (Megastomia) cf. conoidea3
"Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros3 snail
sensu JANSSEN
"Turbonilla (Pyrgolampros) undulata" = Pyrgolampros3 snail
sensu JANSSEN
Turbonilla (Turbonilla) gastaldi3 snail
sensu JANSSEN
Turbonilla (Turbonilla) koeneniana3 snail
sensu JANSSEN
Odostomia (Brachystomia) sp.3 Fleming 1817 snail
 Cerithioidea - Cerithiidae
"Bittium (Bittium) spina" = Bittium spina2
"Bittium (Bittium) spina" = Bittium spina2 Partsch 1856 cerith snail
 Cerithioidea - Turritellidae
Turritella subangulata2
 Pseudomelanoidea - Pseudomelaniidae
sensu lato Inquisitor borealis3
 Vanikoroidea - Eulimidae
Balcis (Balcis) alba2, Eulima (Eulima) bilineata2, Eulima (Eulima) glabra2
 Sorbeoconcha - Vitrinellidae
Circulus praecedens2, Circulus hennei2
Circulus praecedens2 snail
C. praecedens for. praecedens
Circulus hennei2 Glibert 1952 snail
 Sorbeoconcha - Rissoidae
sensu lato Alvania holsatica2, Alvania (Alvania) basisulcata2
 Sorbeoconcha - Triviidae
Erato (Erato) germanica2
Erato (Erato) germanica2 Schilder 1929 snail
 Sorbeoconcha - Cassidae
"Semicassis (Echinophoria) bicoronata" = Echinophoria2, "Semicassis (Semicassis) miolaevigata" = Phalium (Semicassis) miolaevigata2
"Semicassis (Echinophoria) bicoronata" = Echinophoria2 snail
"Semicassis (Semicassis) miolaevigata" = Phalium (Semicassis) miolaevigata2 Sacco 1890 snail
 Neogastropoda - Nassariidae
Nassarius bocholtensis2, Nassarius tenuistriatus2, Nassarius voorthuyseni2, Nassarius karinae2
Nassarius karinae2 Wienrich 2001 snail
 Neogastropoda - Columbellidae
Anachis (Costoanachis) hosiusi2, Anachis (Costoanachis) terebralis2, Mitrella (Macrurella) nassoides2
 Neogastropoda - Fasciolariidae
Streptodictyon sexcostatus2
 Neogastropoda - Buccinidae
Phos (Phos) decussatus2
 Neogastropoda - Cancellariidae
Sveltia varicosa2, Trigonostoma (Trigonostoma) aperta2, Trigonostoma (Trigonostoma) extractrix2, Trigonostoma (Trigonostoma) cf. spinifera2, Cancellaria (Merica) contorta2, "Trigonostoma (Ventrilia) pouwi" = Ventrilia2, Babylonella fusiformis2
Sveltia varicosa2 snail
S. varicosa paucicostata
Cancellaria (Merica) contorta2 snail
C. contorta gelriana
"Trigonostoma (Ventrilia) pouwi" = Ventrilia2 snail
 Neogastropoda - Muricidae
Lyrotyphis (Eotyphis) fistulosus2, Typhis (Typhis) ? pungens2, "Murex (Haustellum) inornatus" = Haustellum2, "Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta2
Typhis (Typhis) ? pungens2 Solander in Brander and Brander 1766 murex snail
"Murex (Haustellum) inornatus" = Haustellum2 murex snail
"Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta2 Bronn 1831 murex snail
 Neogastropoda - Olividae
Oliva (Strephona) dufresnei2
Oliva (Strephona) dufresnei2 Basterot 1825 olive snail
 Neogastropoda - Ancillariidae
"Ancilla (Baryspira) obsoleta" = Amalda obsoleta2
"Ancilla (Baryspira) obsoleta" = Amalda obsoleta2 Brocchi 1814 snail
 Neogastropoda - Borsoniidae
Asthenotoma pannoides3, Asthenotoma pannus3, Bathytoma jugleri3
 Neogastropoda - Raphitomidae
Teretia anceps3, sensu lato Teretia "sp. 2"3, Mioawateria andersoni3, Metuonella grippi3
sensu lato Teretia "sp. 2"3 Norman 1888 snail
 Neogastropoda - Turridae
Fusiturris aquensis3, Fusiturris duchastelii3, Nitidiclavus maitrejus3, Boreodrillia hosiusi3, sensu lato Unedogemmula boreoturricula3, sensu lato Unedogemmula hanseata3, sensu lato Unedogemmula stoffelsii3, Gemmula (Gemmula) denticula3, Gemmula (Gemmula) zimmermanni3
Gemmula (Gemmula) denticula3 turrid
G. denticula borealis
 Neogastropoda - Conidae
"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)3
"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)3 cone shell
 Neogastropoda - Drilliidae
Splendrillia selenkae3
 Neogastropoda - Clavatulidae
Clavatula boreointerrupta3
 Neogastropoda - Mangeliidae
Benthomangelia aff. obtusangula3, sensu lato Mangelia andersoni n. sp.3, Sorgenfreispira "sp. 2"3, Sorgenfreispira tenella3
sensu lato Mangelia andersoni n. sp.3 Wienrich and Janssen 2007 snail
Sorgenfreispira "sp. 2"3 Moroni 1979 snail
Sorgenfreispira tenella3 Conrad 1848 snail
 Neogastropoda - Clathurellidae
Pleurotomoides luisae3, Pleurotomoides simplex3, ? Ceritoturris hannoverana3
 Neogastropoda - Terebridae
Strioterebrum hoernesi3, Strioterebrum ronaldi3
Strioterebrum ronaldi3 auger snail
 Neogastropoda - Costellariidae
Orthosurcula steinvorthi3
 Sorbeoconcha - Xenophoridae
Xenophora deshayesi2
Xenophora deshayesi2 Michelotti 1847 carrier shell
 Sorbeoconcha - Aporrhaidae
Aporrhais (Aporrhais) alata2
 Sorbeoconcha - Naticidae
Naticidae indet.2, Euspira edithae2, Euspira staringi2, Euspira helicina2, Tectonatica miopusilla2, "Natica sensu lato tigrina" = Cochlis tigrina2
Naticidae indet.2 Guilding 1834 moon snail
Naticidae juv. et indet.
Euspira edithae2 moon snail
Euspira staringi2 moon snail
Euspira helicina2 Brocchi 1814 moon snail
E. aggr. helicina
"Natica sensu lato tigrina" = Cochlis tigrina2 Röding 1798 moon snail
 Sorbeoconcha - Triphoridae
Norephora (Norephora) fritschi2
 Sorbeoconcha - Triforidae
Cerithiella (Cerithiella) genei2
 Neotaenioglossa - Calyptraeidae
Calyptraea (Calyptraea) chinensis2
Calyptraea (Calyptraea) chinensis2 Linnaeus 1758 slipper shell
 Epitonioidea - Epitoniidae
"Amaea (Scalina) subreticula" = Scalina2, Cirsotrema (Cirsotrema) crassicostatum2, Opalia (Pliciscala) pertusa2
"Amaea (Scalina) subreticula" = Scalina2 wentletrap
Cirsotrema (Cirsotrema) crassicostatum2 Deshayes 1839 wentletrap
 Opisthobranchiata -
Roxania utriculus3
 Opisthobranchiata - Cylichnidae
Cylichna (Cylichna) pseudoconvoluta3, Scaphander grateloupi3
 Opisthobranchiata - Retusidae
Volvulella (Volvulella) acuminata3, Retusa (Cylichnina) elongata3
1Wienrich 1999; 2Wienrich 2001; 3Wienrich and Janssen 2007