PBDB Collection

Ksar Metlili (KM-B') (Jurassic to of Morocco)

Also known as Ksar Met-Lili

Where: Figuig, Morocco (32.7° N, 3.2° W)

• Paleocoordinates: 16.2° N, 9.7° E (Wright 2013)

• coordinate based on nearby landmark

• small collection-level geographic resolution

When: Ksar Metlili Formation, Late/Upper Tithonian to Late/Upper Tithonian (150.8 - 140.2 Ma)

• original Fm name "Couches Rouges". Localities KM-A1, A2, B', C, D1 and D2 are fossiliferous loci separated by a few metres in the same calcareous fossiliferous sandstone bed.

Environment/lithology: deltaic; lenticular, fine-grained, medium, red, calcareous sandstone

• The Ksar Metlili Formation was likely deposited in a river-dominated deltaic plain, close to the sea. All the vertebrate-bearing beds sampled in the various KM loci show this kind of transitional facies... The KM microvertebrate lens is intercalated between two marine horizons. There is a mixing of continental (wood, mammals, dinosaurs, etc.), brackish (bivalves) and undisputedly marine (echinoderms, coral, brachiopods, etc.) forms in this horizon, which suggests a near-shore, deltaic depositional environment, probably influenced by eustatic variations.

• The sedimentary facies of the Ksar Metlili Formation are similar to those of the continental lower beds of the Anoual Formation (fine to-medium-grained sandstones, overturned cross-stratifications, ripple marks, root traces).

Size classes: macrofossils, mesofossils

Collected by Lasseron et al in 2010

Collection methods: bulk, chemical, sieve

Primary reference: M. Lasseron, R. Allain, E. Gheerbrant, H. Haddoumi, N.-E. Jalil, G. Métais, J.-C. Rage, R. Vullo, and S. Zouhri. 2019. New data on the microvertebrate fauna from the Upper Jurassic or lowest Cretaceous of Ksar Metlili (Anoual Syncline, eastern Morocco). Geological Magazine 1-26 [T. Cleary/T. Cleary]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 204936: authorized by Terri Cleary, entered by Terri Cleary on 27.09.2019

Creative Commons license: CC0 (CC0)

Taxonomic list

• Lasseron et al (2019) mention the new material comprises 24,541 microvertebrate elements. Here I use the number of elements listed for each taxon, which are split between the specimen numbers listed (often 1000+ per specimen number). How these elements are split between specimens can be found in the supp info of Lasseron et al 2019.

Show authors, comments, and common names

Osteichthyes
Therapsida -	cf. Cynodontia "indet. morphotype 1", cf. Cynodontia "indet. morphotype 2" cf. Cynodontia "indet. morphotype 1" Owen 1861 cynodont KM-B'-43-44, 98, single teeth cf. Cynodontia "indet. morphotype 2" Owen 1861 cynodont Not mentioned in supp, possibly KM-B'-99, which is marked as "Cynodontia".
Mammalia
-	Mammalia indet. Mammalia indet. Linnaeus 1758 mammal KM-B'-31-32, both single incisor teeth; 33-34, left dentary; 35, dentary; 38, right dentary; 50, mandibular fragment. Not mentioned in main text, from supp info.
Eutriconodonta -	Eutriconodonta indet. Eutriconodonta indet. Kermack et al. 1973 mammal KM-B'-3, 6, single lower molars; 8, 30, single teeth
Symmetrodonta -	"Symmetrodonta" indet. "Symmetrodonta" indet. symmetrodont KM-B'-36, teeth. Not mentioned in main text, from supp info.
Dryolestoidea -	? Dryolestoidea indet. ? Dryolestoidea indet. mammal "or ?Eutriconodonta". KM-B'-7, tooth. Not mentioned in main text, from supp info.
Dryolestida - Donodontidae	Donodon perscriptoris Donodon perscriptoris mammal KM-B'-2, left lower molar; 11, right upper molar; 95, right upper molar
Dryolestoidea - Dryolestidae	Dryolestidae indet. Dryolestidae indet. Marsh 1879 mammal Probably the same as "Dryolestoidea" in the supplement. KM-B'-1, left lower molar; 37, teeth; 96-97, single left lower molars
Tribosphenida -	Tribotherium africanum, Hypomylos phelizoni, Hypomylos micros Tribotherium africanum Sigogneau-Russell 1991 mammal KM-B'-5, right upper molar Hypomylos phelizoni Sigogneau-Russell 1992 mammal KM-B'-4, left lower molar; 9-10, single right lower molars Hypomylos micros Sigogneau-Russell 1995 mammal [no mention of specimen no.s in supp info]
Osteichthyes
Sauropsida -	Sauropsida indet. Sauropsida indet. Huxley 1864 amniote KM-B'-87, teeth
Reptilia
Loricata -	Crocodylomorpha indet. Crocodylomorpha indet. Hay 1930 crocodilian KM-B'-40, osteoderms; KM-B'-45, mandibular fragment; 46, 52, 91, all teeth
Loricata - Atoposauridae	Atoposauridae indet., cf. Theriosuchus indet. Atoposauridae indet. Gervais 1871 crocodilian KM-B'-47, 48, teeth cf. Theriosuchus indet. Owen 1878 crocodilian KM-B'-42, premaxillary
Loricata - Teleosauridae	Teleosauridae indet. Teleosauridae indet. Geoffroy 1831 marine crocodile KM-B'-51, teeth; KM-B'-53, tooth
Pterosauria -	Pterosauria indet. Pterosauria indet. pterosaur KM-B'-54-55, teeth
Theropoda -	Theropoda indet., Maniraptora indet. Theropoda indet. Marsh 1881 theropod KM-B'-21, claw; KM-B'-23-25, teeth; 94, claw (ungual?) Maniraptora indet. Gauthier 1986 maniraptoran KM-B'-22, tooth. Not mentioned in main text, from supp info.
Theropoda - Dromaeosauridae	Dromaeosauridae indet. Dromaeosauridae indet. Colbert and Russell 1969 maniraptoran KM-B'-26, teeth. Not mentioned in main text, from supp info.
Choristodera -	Choristodera indet. Choristodera indet. Cope 1876 choristodere KM-B'-49, dorsal vertebrae
Eosuchia -	Sphenodontia indet. Sphenodontia indet. lepidosaur KM-B'-14, dorsal vertebra
Testudinata -	Testudinata indet. Testudinata indet. Oppel 1811 turtle KM-B'-39, 41, osteoderms
Squamata -	Squamata indet., Scincomorpha indet., Tarratosaurus anoualensis Squamata indet. squamates KM-B'-28, dorsal vertebrae; 29, jaw fragment; 90, 92, single teeth Scincomorpha indet. Camp 1923 squamates KM-B'-19, right dentary Tarratosaurus anoualensis Broschinksi and Sigogneau-Russell 1996 lizard KM-B'-20, right dentary. Not mentioned in main text, from supp info.
Amphibia
Temnospondyli - Albanerpetontidae	Anoualerpeton unicus Anoualerpeton unicus Gardner et al. 2003 tetrapod KM-B'-12-13, left dentary
Salientia -	Anura indet. Anura indet. Fischer von Waldheim 1813 frog KM-B'-15, right ilium; 16, scapula; 17, dorsal vertebrae; 27, dorsal vertebrae of a hatchling or neotenic; 86, long bone fragments
Salientia - Alytidae	cf. Alytidae indet. cf. Alytidae indet. Fitzinger 1843 midwife toads KM-B'-18, dorsal vertebrae
Osteichthyes
Dipnoi - Ptychoceratodontidae	cf. Ptychoceratodontidae indet. cf. Ptychoceratodontidae indet. Martin 1982 lungfish KM-B'-56, teeth
Actinopterygii -	Actinopterygii indet. Actinopterygii indet. Cope 1887 KM-B'-62, 73-74, 80, scales; KM-B'-63, 67-68, 76-77, 79, all fin rays; 69, vertebrae; 81, mandibular fragments. Not mentioned in main text, from supp info.
Actinopteri
Teleostei -	? Siluriformes indet., Osteoglossiformes indet. ? Siluriformes indet. catfish "or ?Polypteriformes". KM-B'-57, fin spine Osteoglossiformes indet. Berg 1940 KM-B'-89, teeth
Ginglymodi -	Ginglymodi indet. Ginglymodi indet. Cope 1871 KM-B'-58 and 60, 66, all ichthyoliths; KM-B'-59, 61, 64-65, 72, 84, all scales
Chondrichthyes
Hybodontiformes - Hybodontidae	Egertonodus basanus Egertonodus basanus Egerton 1845 elasmobranch